Atriplex patula
Atriplex suckleyi
common orache, halberd-leaf orache, spear orach, spear orache, spear oracle, spear saltbush, spear saltweed, spearscal e, spearscale orache
Suckley's orach
mostly erect and branched, branches green, obtusely angled or striate, glabrate.
terete, often tinged with red, sparsely mealy or glabrous.
alternate except the proximalmost, petiolate;
blade green on both sides, rhombic-lanceolate to lanceolate, oblong, or narrowly lance-oblong or hastate-ovate, 25–120 × 3–40(–75) mm, entire or toothed, proximal ones broadly cuneate or sometimes hastate subbasally with obliquely antrorse basal lobes, distal cauline leaves lanceolate and entire.
numerous, alternate, sessile;
blade lanceolate to elliptic or less commonly ovate, 7–35 × (2–)4–10(–11) mm, thick and succulent, base acute, margin entire, apex acute or acuminate, glaucous, sparsely farinose when young.
compact or interrupted spiciform or paniculiform clusters.
mostly 5-merous.
in small glomerules, these in distal axils or in short, dense or interrupted, mostly simple, terminal spikes, perianth cup-shaped, frequently pinkish, lobes each with fleshy crest.
solitary or few in leaf axils, calyx within bracteoles of 3 or 4, entire or lobed, distinct, obtuse, hyaline sepals.
dimorphic: brown, 2.5–3(–3.5) mm wide, or black, 1–2 mm wide;
radicle of brown seeds subbasal to median, antrorse.
brown, 1.5 × 1.2 mm.
bracteoles green becoming black, rhombic to rhombic-triangular, or ovate-rhombic, compressed, ± uniformly sized, 2–7(–20) mm, base mostly hastate, acute, margin united almost to middle, entire or sparingly toothed, surfaces tuberculate.
bracteoles small and difficult to find, sessile, ovate, 2 × 1.5 mm, membranous, united to apex, without appendages, scurfy, each pistil subtended by a perianth.
= 36.
Atriplex patula
Atriplex suckleyi
Atriplex patula appears to have been a rather recent introduction in North America from Eurasia, not arriving perhaps until sometime in the early to mid-eighteenth century. It simulates depauperate specimens of A. dioica, A. glabriuscula, and other similar species when leaves are reduced to a near-linear profile. Such specimens are difficult if not impossible to assign to any of the species.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
At specific rank, the epithet suckleyi clearly has priority. Stutz et al. (1993) resurrected Endolepis Torrey and placed within it two completely disparate species, E. dioica and E. covillei Standley [Atriplex covillei (Standley) J. F. Macbride]. The interpretation of the genus by Stutz et al. stands on the presence of perianth segments subtending the ovary within the fruiting bracteoles, lack of Kranz leaf anatomy, and other more equivocal characteristics. The shared features hardly indicate near affinity, however. The two taxa are otherwise grossly dissimilar. Fundamentally, the genus Endolepis as resurrected by H. C. Stutz et al. stands on the basis of a single character: the presence of perianth segments. Kranz anatomy rises and falls, both within subg. Atriplex and subg. Obione. Thus, coincidence of the non-Kranz criterion is subject to interpretation. Perianth segments subtending the ovary within the enclosing bracteoles, per se, appear to be of independent origin. And, the peculiar nature of the staminate calyx in A. suckleyi (a major determining condition in establishment of the genus Endolepis by Torrey) is not present in A. covillei. Certainly the two species included within Endolepis appear to be as closely allied to other species of Atriplex as they are to each other.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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