Atriplex patula
Atriplex californica
common orache, halberd-leaf orache, spear orach, spear orache, spear oracle, spear saltbush, spear saltweed, spearscal e, spearscale orache
California orach, California saltbush
mostly erect and branched, branches green, obtusely angled or striate, glabrate.
many branched, subterete, 1.5–5 dm, white scurfy when young.
alternate except the proximalmost, petiolate;
blade green on both sides, rhombic-lanceolate to lanceolate, oblong, or narrowly lance-oblong or hastate-ovate, 25–120 × 3–40(–75) mm, entire or toothed, proximal ones broadly cuneate or sometimes hastate subbasally with obliquely antrorse basal lobes, distal cauline leaves lanceolate and entire.
numerous, often crowded, alternate or proximalmost opposite;
blade narrowly lanceolate to narrowly oblanceolate or elliptic, 4–20 × 1–5 mm, acute at both ends, gray scurfy.
compact or interrupted spiciform or paniculiform clusters.
mostly 5-merous.
in terminal bracteate spikes, or mixed with pistillate in rather dense axillary clusters, 4-merous.
dimorphic: brown, 2.5–3(–3.5) mm wide, or black, 1–2 mm wide;
radicle of brown seeds subbasal to median, antrorse.
dark (black), 1–2 mm.
bracteoles green becoming black, rhombic to rhombic-triangular, or ovate-rhombic, compressed, ± uniformly sized, 2–7(–20) mm, base mostly hastate, acute, margin united almost to middle, entire or sparingly toothed, surfaces tuberculate.
bracteoles sessile, rhombic-ovate to ovate, scarcely united, 2.5–4 mm, margin entire, acute.
= 36.
Atriplex patula
Atriplex californica
Atriplex patula appears to have been a rather recent introduction in North America from Eurasia, not arriving perhaps until sometime in the early to mid-eighteenth century. It simulates depauperate specimens of A. dioica, A. glabriuscula, and other similar species when leaves are reduced to a near-linear profile. Such specimens are difficult if not impossible to assign to any of the species.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
H. M. Hall and F. E. Clements (1923) placed great emphasis on the inferior radicle, dioecious habit, and free bracts in stating that there are no close relatives of Atriplex californica in North America. P. C. Standley (1916) regarded the radicle as lateral or superior, not inferior, and placed the species at the beginning of his treatment of the American species. Plants of A. californica, however, form a mirror-image, matched pair with A. watsonii, with which they share habit, leaf conformation, staminate glomerules arranged, at least partially, in terminal interrupted spikes, and Kranz anatomy, but from which they differ in the radicle position, valves of the fruiting bracteoles free beyond the middle, monoecious habit, and mostly alternate leaves. The interpretation by Hall and Clements of radicle position as fundamental is made problematic by the apparent random occurrence of a great many morphologic features from place to place within the genus and often within the taxon. Each character fails individually as having definitive taxonomic importance, making difficult or impossible an ultimately satisfactory phylogenetic arrangement. It is not, however, the character that makes the species, rather, it is the entire syndrome of features that constitutes the taxon. Most certainly A. californica is more closely allied to American taxa than to those from other regions of the world.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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