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Parish's brittlescale, Parish's saltbush, Parrish's brittlescale

allscale, allscale saltbush, cattle saltbush, cattle-spinach, desert saltbush

Habit Herbs, erect or spreading to prostrate, 0.5–3 dm; branches almost horizontal to ascending, fragile, white scurfy or villous (in var. parishii). Shrubs, dioecious or sub-monoecious, mainly 10–20 dm.
Leaves

numerous, all or nearly all opposite or almost all alternate, distal ones imbricate or widely separated, tending to recurve;

blade lanceolate to ovate, (2–)4–10 × 3–8 mm, rigid, base mostly rounded to cordate, margin entire, gray to white, densely scurfy (or hairy).

tardily deciduous, alternate, sessile or nearly so;

blade spatulate to obovate or oblong, 3–15(–2.) × 2–4 mm, margin entire, apex typically acute.

Staminate flowers

mostly in distal axils pistillate in proximal axils, or mostly in terminal spike (var. persistens), or partly so (var. subtilis).

appearing silvery, in clusters 1.5–3 mm wide, borne in paniculate, naked spikes 5–25 cm;

anthers yellow.

Pistillate flowers

with fruiting bracteoles sessile, cuneate-orbicular to semicircular, 1.5–2.5 × 2–3 mm, united to near middle, margin deeply laciniate-dentate, tuberculate or sometimes smooth dorsally.

Seeds

dark brown or almost black, 0.8–1.5 mm.

pale brown, 1–1.5 mm.

Fruiting

bracteoles sessile, ovate or rhombic, slightly compressed to thickened, 2–3.5(–4) mm and about as broad or sometimes broader, often subhastately lobed, united 1/2 of length, entire or with few teeth on each side, tuberculate on 1 or both faces.

2n

= 18, 36.

Atriplex parishii

Atriplex polycarpa

Phenology Flowering spring–fall.
Habitat Warm desert shrub communities (creosote bush, ambrosia, shadscale, mesquite, saltgrass, etc.), mainly in fine-textured saline substrates
Elevation 60-1500 m (200-4900 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NV; UT; n Mexico (Baja California to Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 5 (5 in the flora).

The Atriplex parishii complex consists of a series of microphyllous, low clump-forming annuals apparently disjunct from each other in the Central Valley of California and in near coastal southern California. Often they occupy vernal pools that dry as the season progresses; the substrates in all cases evidently are saline or alkaline, or both. For the most part, the bracteate distal leaves are cordate to rounded at the base, and spreading to spreading-ascending, and the fruiting bracteoles are mainly less than 3.5 mm in length.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Atriplex polycarpa consists of two or more chromosomal races based on different polyploid levels. The races form hybrids with Atriplex canescens in south California, resulting in partially stabilized entities known as varieties laciniata and macilenta. The plants evidently form hybrids with A. lentiformis.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruiting bracteoles persistent, ± thickened and/or tuberculate on 1 or both faces
→ 2
1. Fruiting bracteoles deciduous, compressed, faces smooth or tuberculate
→ 3
2. Stems, even in inflorescences, mainly over 1.5 mm thick; staminate flowers borne mainly in terminal spikes; fruiting bracteoles 2.5-3.5(-4) mm; Colusa, Glenn, Merced, Solano, Stanislaus, and Tulare counties.
var. persistens
2. Stems all or at least above, filiform, less than 1.5 mm thick; staminate flowers borne mainly in axillary glomerules, only sometimes somewhat spicate; fruiting bracteoles 2.4-2.8 mm; Fresno, Kings, Kern, Madera, and Tulare counties
var. subtilis
3. Plants with elongate slender hairs in inflorescences; foliose bracts of inflorescence spreading, tips ± recurved; plants of Los Angeles (or Orange), Riverside, and San Diego counties
var. parishii
3. Plants merely scurfy in inflorescences; foliose bracts of inflorescence spreading-ascending, tips not or not especially recurved
→ 4
4. Leaves all or mainly opposite, 3-7 mm; fruiting bracteoles with appendages not in rows, on abaxial surface only; Glenn, Solano, and Yolo counties
var. depressa
4. Leaves all or mainly alternate, 4-10 mm; fruiting bracteoles with faces smooth; Tulare and Kern counties
var. minuscula
Source FNA vol. 4, p. 356. FNA vol. 4, p. 376.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Pusillae Chenopodiaceae > Atriplex > subg. Pterochiton
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. parishii var. depressa, A. parishii var. minuscula, A. parishii var. parishii, A. parishii var. persistens, A. parishii var. subtilis
Synonyms Obione parishii Obione polycarpa, A. curvidensB
Name authority S. Watson: Proc. Amer. Acad. Arts 17: 377. (1882) (Torrey) S. Watson: Proc. Amer. Acad. Arts 9: 117. (1874)
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