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Parish's brittlescale, Parish's saltbush, Parrish's brittlescale

Kleberg orach, Kleberg's saltbush

Habit Herbs, erect or spreading to prostrate, 0.5–3 dm; branches almost horizontal to ascending, fragile, white scurfy or villous (in var. parishii). Herbs, with ligneous vertical taproot 5–9 mm thick; bark pale.
Stems

erect, diffuse;

branches alternate, numerous, horizontal or distally ascending, terete, 1.5–4 dm, densely white farinose when young, glabrate in age;

bark becoming pale brownish white and flaky;

internodes mostly shorter than 1(–2.4) cm.

Leaves

numerous, all or nearly all opposite or almost all alternate, distal ones imbricate or widely separated, tending to recurve;

blade lanceolate to ovate, (2–)4–10 × 3–8 mm, rigid, base mostly rounded to cordate, margin entire, gray to white, densely scurfy (or hairy).

alternate, proximalmost subopposite, sessile;

blade ovate-deltoid, 0.5–1.5(–2.5) × 5–12(–15) mm, rather firm and flat, base rounded, truncate, or slightly clasping, margin entire or toothed, apex acute, very densely canescent-farinose with a greenish yellow tinge.

Flowers

sessile, axillary, inconspicuous, mostly in leafy lateral branches with extremely short internodes, arising toward tips of secondary branches.

Staminate flowers

mostly in distal axils pistillate in proximal axils, or mostly in terminal spike (var. persistens), or partly so (var. subtilis).

in most distal axils, 2 mm wide;

sepals 3–5, mostly hyaline, curved elliptic, 1.5–1.7 mm, mucronulate, farinose dorsally.

Pistillate flowers

densely farinose;

bracteoles adnate to ovary.

Seeds

dark brown or almost black, 0.8–1.5 mm.

dark reddish brown, round-lenticular, 1.5 mm wide, shining;

radicle superior.

Fruiting

bracteoles sessile, ovate or rhombic, slightly compressed to thickened, 2–3.5(–4) mm and about as broad or sometimes broader, often subhastately lobed, united 1/2 of length, entire or with few teeth on each side, tuberculate on 1 or both faces.

bracteoles variably and irregularly 3–7-cleft, ovate-orbicular, 3.1–4.7 × (3.2–)4–7 mm, typically somewhat constricted below middle, with terminal lobes (1–)1.5–2.8 mm, densely scurfy, faces doubly cristate or smooth.

Atriplex parishii

Atriplex klebergorum

Phenology Flowering summer.
Habitat In silty or clay loam soils
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; of conservation concern; near sea level
[BONAP county map]
Discussion

Varieties 5 (5 in the flora).

The Atriplex parishii complex consists of a series of microphyllous, low clump-forming annuals apparently disjunct from each other in the Central Valley of California and in near coastal southern California. Often they occupy vernal pools that dry as the season progresses; the substrates in all cases evidently are saline or alkaline, or both. For the most part, the bracteate distal leaves are cordate to rounded at the base, and spreading to spreading-ascending, and the fruiting bracteoles are mainly less than 3.5 mm in length.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

The species was noted by its author as apparently belonging in sect. Argenteae Standley; I concur with that alignment. The overall shape of the bracteoles with a subterminal constriction is reminiscent of those of Atriplex powellii, but the bracteole shape is otherwise distinctive, and the long marginal teeth and occasional elongate cristate processes on the faces are unmatched elsewhere in the Argenteae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruiting bracteoles persistent, ± thickened and/or tuberculate on 1 or both faces
→ 2
1. Fruiting bracteoles deciduous, compressed, faces smooth or tuberculate
→ 3
2. Stems, even in inflorescences, mainly over 1.5 mm thick; staminate flowers borne mainly in terminal spikes; fruiting bracteoles 2.5-3.5(-4) mm; Colusa, Glenn, Merced, Solano, Stanislaus, and Tulare counties.
var. persistens
2. Stems all or at least above, filiform, less than 1.5 mm thick; staminate flowers borne mainly in axillary glomerules, only sometimes somewhat spicate; fruiting bracteoles 2.4-2.8 mm; Fresno, Kings, Kern, Madera, and Tulare counties
var. subtilis
3. Plants with elongate slender hairs in inflorescences; foliose bracts of inflorescence spreading, tips ± recurved; plants of Los Angeles (or Orange), Riverside, and San Diego counties
var. parishii
3. Plants merely scurfy in inflorescences; foliose bracts of inflorescence spreading-ascending, tips not or not especially recurved
→ 4
4. Leaves all or mainly opposite, 3-7 mm; fruiting bracteoles with appendages not in rows, on abaxial surface only; Glenn, Solano, and Yolo counties
var. depressa
4. Leaves all or mainly alternate, 4-10 mm; fruiting bracteoles with faces smooth; Tulare and Kern counties
var. minuscula
Source FNA vol. 4, p. 356. FNA vol. 4, p. 353.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Pusillae Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Argenteae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. parishii var. depressa, A. parishii var. minuscula, A. parishii var. parishii, A. parishii var. persistens, A. parishii var. subtilis
Synonyms Obione parishii
Name authority S. Watson: Proc. Amer. Acad. Arts 17: 377. (1882) M. C. Johnston: SouthW. Naturalist 6: 49. (1961)
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