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broadscale, mound saltbush, New Mexico saltbush, silver saltbush

Coville's orach

Habit Subshrubs, dioecious, clump forming, mainly 2–8 dm and as wide, woody at base. Herbs, spreading, 1–4(–5) dm and as broad.
Stems

stiffly erect;

branchlets terete.

terete, sparsely scurfy when young.

Leaves

tardily deciduous, alternate or proximal-most subopposite, shortly petiolate;

blade gray green, oblong-ovate to elliptic or orbiculate, 8–30(–35) × 6–20 mm, margin entire or rarely dentate, apex rounded to retuse or obtuse.

petiole to 1/2 as long as blade (becoming subsessile distally);

blade green or finally stramineous, (10–)20–50 × 6–30 mm, firm, base abruptly acute to narrowly cuneate, apex acute to attenuate, sparsely scurfy.

Staminate flowers

yellow, in clusters 2–3 mm wide, borne in panicles 6–30 cm.

in sessile glomerules in distal axils, often mixed with pistillate ones, staminate calyx deeply 5-cleft;

lobes obtuse, not appendaged.

Pistillate flowers

in small, very numerous glomerules in axils of elongated, terminal leafy-bracteate spikes or finally paniculate.

with calyx of (1–)3(–5) hyaline sepals.

Seeds

brown, 2.4–2.8 mm.

dark reddish brown, 1–1.5 mm.

Fruiting

bracteoles sessile or substipitate, 4–5 × 5–9 mm, base broadly cuneate, margin sharply toothed, apical tooth subtended by 2–6 equal or smaller teeth, faces smooth or rarely tuberculate.

bracteoles sessile or stipitate, 6–12 × 4–7 mm, margin mostly 3-lobed, with elongate terminal lobe triangular to lanceolate, 2 short rounded lobes at base or sides merely rounded at base, united to beyond middle.

Atriplex obovata

Atriplex covillei

Phenology Flowering summer–fall. Flowering summer–fall.
Habitat Fine-textured substrates, with salt desert shrub and lower pinyon-juniper communities Mixed saltbush-greasewood, rabbitbrush, warm desert shrub, and salt grass communities in saline substrates
Elevation 1500-2000 m (4900-6600 ft) 800-1700 m (2600-5600 ft)
Distribution
from FNA
AZ; CO; NM; TX; UT; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
Discussion

Endolepis covillei was treated within the synonymy of Atriplex phyllostegia (Torrey) S. Watson by H. M. Hall and F. E. Clements (1923).

H. C. Stutz et al. (1993) placed Atriplex covillei within Endolepis, based in large part on the presence of a perianth subtending the pistil within the fruiting bracteoles and on the lack of Kranz anatomy in the leaves. The pattern of venation is, nevertheless, very similar to that in species with Kranz anatomy. The presence of perianth scales in the pistillate flowers of A. covillei has been regarded as evidence of relationship with A. suckleyi. Despite placement of these taxa within Endolepis by Stutz et al. Atriplex covillei is possibly more closely allied to the morphologically similar and partially sympatric A. phyllostegia than it is to strongly dissimilar and the distantly disjunct A. suckleyi. Stutz and his associates placed great emphasis on the presence of reduced perianth segments subtending the pistil within the fruiting bracteoles of A. covillei. Calyces per se, otherwise known only in A. suckleyi and A. pleiantha, probably have arisen independently. Their presence does not necessarily indicate a close relationship. Stutz et al. pointed to other differences aside from the calyx of the pistillate flowers, and it is apparent that the two entities can stand as distinct species. To segregate A. covillei within a separate genus and to ally it with a species to which its relationships are obscure at best, stretches logic beyond reason.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 371. FNA vol. 4.
Parent taxa Chenopodiaceae > Atriplex > subg. Pterochiton Chenopodiaceae > Atriplex > subg. Obione > sect. Covilleiae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Synonyms A. greggii, A. jonesii, A. obovata var. tuberata Endolepis covillei
Name authority Moquin-Tandon: Chenop. Monogr. Enum., 61. (1840) (Standley) J. F. Macbride: Contr. Gray Herb. 53: 11. (1918)
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