Atriplex obovata |
Atriplex californica |
|
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broadscale, mound saltbush, New Mexico saltbush, silver saltbush |
California orach, California saltbush |
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Habit | Subshrubs, dioecious, clump forming, mainly 2–8 dm and as wide, woody at base. | Herbs, monoecious or dioecious, prostrate or procumbent-decumbent, from fleshy fusiform or variously shaped taproot. |
Stems | stiffly erect; branchlets terete. |
many branched, subterete, 1.5–5 dm, white scurfy when young. |
Leaves | tardily deciduous, alternate or proximal-most subopposite, shortly petiolate; blade gray green, oblong-ovate to elliptic or orbiculate, 8–30(–35) × 6–20 mm, margin entire or rarely dentate, apex rounded to retuse or obtuse. |
numerous, often crowded, alternate or proximalmost opposite; blade narrowly lanceolate to narrowly oblanceolate or elliptic, 4–20 × 1–5 mm, acute at both ends, gray scurfy. |
Staminate flowers | yellow, in clusters 2–3 mm wide, borne in panicles 6–30 cm. |
in terminal bracteate spikes, or mixed with pistillate in rather dense axillary clusters, 4-merous. |
Pistillate flowers | in small, very numerous glomerules in axils of elongated, terminal leafy-bracteate spikes or finally paniculate. |
|
Seeds | brown, 2.4–2.8 mm. |
dark (black), 1–2 mm. |
Fruiting | bracteoles sessile or substipitate, 4–5 × 5–9 mm, base broadly cuneate, margin sharply toothed, apical tooth subtended by 2–6 equal or smaller teeth, faces smooth or rarely tuberculate. |
bracteoles sessile, rhombic-ovate to ovate, scarcely united, 2.5–4 mm, margin entire, acute. |
Atriplex obovata |
Atriplex californica |
|
Phenology | Flowering summer–fall. | Flowering Apr–Nov. |
Habitat | Fine-textured substrates, with salt desert shrub and lower pinyon-juniper communities | Sea bluffs, sandy coasts, crevices in sea cliffs, coastal strands, edges of coastal salt marsh, coastal sage scrub |
Elevation | 1500-2000 m (4900-6600 ft) | 0-50 m (0-200 ft) |
Distribution |
AZ; CO; NM; TX; UT; Mexico
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CA; Mexico
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Discussion | H. M. Hall and F. E. Clements (1923) placed great emphasis on the inferior radicle, dioecious habit, and free bracts in stating that there are no close relatives of Atriplex californica in North America. P. C. Standley (1916) regarded the radicle as lateral or superior, not inferior, and placed the species at the beginning of his treatment of the American species. Plants of A. californica, however, form a mirror-image, matched pair with A. watsonii, with which they share habit, leaf conformation, staminate glomerules arranged, at least partially, in terminal interrupted spikes, and Kranz anatomy, but from which they differ in the radicle position, valves of the fruiting bracteoles free beyond the middle, monoecious habit, and mostly alternate leaves. The interpretation by Hall and Clements of radicle position as fundamental is made problematic by the apparent random occurrence of a great many morphologic features from place to place within the genus and often within the taxon. Each character fails individually as having definitive taxonomic importance, making difficult or impossible an ultimately satisfactory phylogenetic arrangement. It is not, however, the character that makes the species, rather, it is the entire syndrome of features that constitutes the taxon. Most certainly A. californica is more closely allied to American taxa than to those from other regions of the world. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4, p. 371. | FNA vol. 4, p. 366. |
Parent taxa | Chenopodiaceae > Atriplex > subg. Pterochiton | Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Californicae |
Sibling taxa | ||
Synonyms | A. greggii, A. jonesii, A. obovata var. tuberata | |
Name authority | Moquin-Tandon: Chenop. Monogr. Enum., 61. (1840) | Moquin-Tandon: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 13(2): 98. (1849) |
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