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quelite

sack saltbush, sack saltweed, stalk orach, stalk orache

Habit Herbs, annual. Herbs, erect, forming rounded clumps.
Stems

erect, ascending, or procumbent, much branched, obtusely angled, 1–6 dm, stout, scurfy when young.

mainly 0.5–4(–5) dm;

stems usually branched from base, terete, stout, angled;

herbage scurfy.

Leaves

alternate or proximalmost opposite or subopposite;

petiole to 1.5 cm or sessile;

blade paler abaxially, oblong or oval, lanceolate, or elliptic to broadly obovate or narrowly oblong, 10–40(–60) × (2–)4–20 mm, base rounded to cuneate, margin entire or undulate, rarely with 1–2 teeth, apex rounded to acute, mucronate, thin, often densely white scurfy beneath, grayish green and glabrate above.

alternate or proximalmost subopposite, short petiolate or distal ones sessile;

petiole 2–5 mm;

blade cordate-ovate or subreniform to ovate to deltoid-ovate or oval, mainly 6–40 × 4–30 mm, base truncate to subcordate or broadly cuneate, margin entire or in some subhastately lobed or sometimes undulate-dentate, apex acute to rounded.

Pistillate flowers

fascicled in axils.

usually in fascicles of 1–3 in proximal axils.

Seeds

reddish brown, 2 mm.

brownish to whitish, 1.5–2.3 mm.

Staminate

glomerules terminal or in dense or interrupted, terminal or axillary, naked spikes or shortly branched panicles.

glomerules in distal axils or in short, naked, terminal (early deciduous) panicles;

flowers with 5-parted calyx.

Fruiting

bracteoles subsessile or with stipes to 1 mm, cuneate-orbiculate, compressed, 4.5–7 × 3.5–5.6 mm, typically longer than wide, united to middle, apex rounded, 3–5-toothed, teeth subequal, sides irregularly tuberculate or with 2 lateral dentate crests, rarely not appendaged.

bracteoles monomorphic or dimorphic, larger ones on stipes (2–)4–8(–15) mm, others sessile, united at base, round-triangular or suborbicular, 4–6 mm, irregularly and coarsely dentate and with ± densely beset with flat, cristate, or hornlike appendages, smaller bracteoles (lacking or rare in var. asterocarpa) in same axils, oblong to cuneate, 3–4 mm, apex truncate, dentate only at summit, faces smooth.

2n

= 18.

= 18.

Atriplex mucronata

Atriplex saccaria

Phenology Flowering summer–fall.
Habitat Sandy seashores, salt marshes
Elevation 0 m (0 ft)
Distribution
from FNA
AL; FL; LA; MA; MD; MS; NC; NH; NJ; TX; VA
[BONAP county map]
from FNA
AZ; CO; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Problems with prior applications of the name Atriplex mucronata lie in the mistaken determination of the place of publication as Amer. Monthly Mag. & Crit. Rev. 2: 176. 1818 (where the name is only mentioned), instead of 2: 119. 1817 (where accompanied by a description and notes). H. M. Hall and F. E. Clements (1923) evidently relied on P. C. Standley’s (1916) interpretation, which indicated the wrong citation (see also A. dioica Rafinesque as an identical example). Hall and Clements applied the name mucronata to their interpretation of A. patula subsp. hastata or to A. hastata (i.e., to A. prostrata according to this treatment).

The treatment of Obione by C. H. B. A. Moquin-Tandon (1849) included “A. mucronata Rafin.!” as a synonym of O. arenaria. Possibly a sheet in the Prodromus herbarium at Geneva was the basis for that decision. It has two mounted specimens, one labeled A. arenaria Nuttall, collected by Nuttall in “N. Jersey, 1826,” and a second labeled “Atriplex mucronata Rafinesque (A. arenaria Nuttall, N. Jersey) Maritime NY, Rafinesque 1819.” From that information (although the year is 1819, not 1817), and from the description of the taxon, it seems clear that A. arenaria Nuttall is a later synonym of A. mucronata Rafinesque by at least half a year.

H. A. Gleason and A. Cronquist (1991) indicated that this taxon, by whatever name, is “perhaps better treated as a variety of the more tropical Atriplex pentandra (Jacquin) Standley, but the proper nomenclatural combination not yet made.” The present writer agrees with that conclusion, but such subjugation might indicate further contraction into the species of additional closely related taxa, e.g., A. wrightii, which is clearly closely allied as well.

Plants from the coastal states from New England south to New Jersey are much alike and seldom, if ever, display prominent, terminal, naked spikes or panicles with beadlike glomerules of staminate flowers. Plants from Florida westward sometimes have such staminate spikes or panicles.

Specimens from Florida and some from Texas can be distinguished from Atriplex pentandra only with difficulty, especially those individuals with entire leaves. However, the fruiting bracteole length-width ratio and overall shape, with some allowance for overlap, can serve to distinguish most specimens; those of A. mucronata are proportionately longer than broad and, on average, larger.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaves mainly ovate to cordate in profile, the distalmost bracteate ones (and sometimes all) often broadly so in age; fruiting bracteoles with stipes to 10 mm, but seldom exceeding 6 mm; Arizona, New Mexico, Wyoming, s to Texas
var. saccaria
1. Leaves oval to rhombic, the bases truncate to cuneate-attenuate; fruiting bracteoles with stipes of various length; sw Utah and adjacent states
→ 2
2. Leaves oval to suborbiculate or less commonly elliptic; fruiting bracteoles with hornlike appendages, borne on stipes mainly 2-6 mm
var. asterocarpa
2. Leaves rhombic to oval or triangular, base cuneate-attenuate; fruiting bracteoles with flattened processes, borne on stipes mainly 2-15 mm (sessile in some)
var. cornuta
Source FNA vol. 4, p. 362. FNA vol. 4.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Saccariae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. saccaria var. asterocarpa, A. saccaria var. cornuta, A. saccaria var. saccaria
Synonyms A. arenaria, A. cristata var. arenaria, A. pentandra subsp. arenaria A. truncata var. saccaria, Obione saccaria
Name authority Rafinesque: Amer. Monthly Mag. & Crit. Rev. 2(2): 119. (1817) S. Watson: Proc. Amer. Acad. Arts 9: 112. (1874)
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