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quelite

red orach, red orache, redscale, tumbling orach, tumbling orache, tumbling saltweed

Habit Herbs, annual. Herbs, erect, coarse, 1–10(–20) dm.
Stems

erect, ascending, or procumbent, much branched, obtusely angled, 1–6 dm, stout, scurfy when young.

simple or more commonly divaricately branching throughout, branches terete;

herbage whitish scurfy to glabrate.

Leaves

alternate or proximalmost opposite or subopposite;

petiole to 1.5 cm or sessile;

blade paler abaxially, oblong or oval, lanceolate, or elliptic to broadly obovate or narrowly oblong, 10–40(–60) × (2–)4–20 mm, base rounded to cuneate, margin entire or undulate, rarely with 1–2 teeth, apex rounded to acute, mucronate, thin, often densely white scurfy beneath, grayish green and glabrate above.

alternate, short petiolate, blade prominently 3-veined, ovate to lanceolate, mainly 12–80 × 6–50 mm, margin irregularly sinuate-dentate and often subhastately lobed or rarely some entire, apex acute to obtuse.

Flowers

in axillary glomerules or interrupted terminal spikes.

Staminate flowers

with 4 or 5 sepals.

Pistillate flowers

fascicled in axils.

in axillary glomerules of 5–10.

Seeds

reddish brown, 2 mm.

dimorphic: brown, 2–2.5 mm wide, or black, 1–2 mm wide;

radicle inferior.

Staminate

glomerules terminal or in dense or interrupted, terminal or axillary, naked spikes or shortly branched panicles.

Fruiting

bracteoles subsessile or with stipes to 1 mm, cuneate-orbiculate, compressed, 4.5–7 × 3.5–5.6 mm, typically longer than wide, united to middle, apex rounded, 3–5-toothed, teeth subequal, sides irregularly tuberculate or with 2 lateral dentate crests, rarely not appendaged.

bracteoles prominently 3–5-veined, sessile or short stipitate, (3–)4–6(–10) mm and as wide, sometimes subhastately lobed at base, conspicuously dentate, sharply tuberculate to almost smooth on faces.

2n

= 18.

= 18.

Atriplex mucronata

Atriplex rosea

Phenology Flowering summer–fall. Flowering mid summer–fall.
Habitat Sandy seashores, salt marshes Disturbed sites, often in riparian habitats or in barnyards or on animal bed grounds, along roadsides and irrigation canals, with juniper, sagebrush, rabbitbrush, pinyon-juniper, Salsola, Chrysothamnus, Atriplex spp., and other weedy species
Elevation 0 m (0 ft) 0-2600 m (0-8500 ft)
Distribution
from FNA
AL; FL; LA; MA; MD; MS; NC; NH; NJ; TX; VA
[BONAP county map]
from FNA
AZ; CA; CO; FL; ID; MA; MI; MO; MT; ND; NE; NJ; NM; NV; NY; OH; OR; PA; UT; WA; WI; WY; AB; BC; NS; ON; SK; Eurasia [Introduced in North America]
[WildflowerSearch map]
[BONAP county map]
Discussion

Problems with prior applications of the name Atriplex mucronata lie in the mistaken determination of the place of publication as Amer. Monthly Mag. & Crit. Rev. 2: 176. 1818 (where the name is only mentioned), instead of 2: 119. 1817 (where accompanied by a description and notes). H. M. Hall and F. E. Clements (1923) evidently relied on P. C. Standley’s (1916) interpretation, which indicated the wrong citation (see also A. dioica Rafinesque as an identical example). Hall and Clements applied the name mucronata to their interpretation of A. patula subsp. hastata or to A. hastata (i.e., to A. prostrata according to this treatment).

The treatment of Obione by C. H. B. A. Moquin-Tandon (1849) included “A. mucronata Rafin.!” as a synonym of O. arenaria. Possibly a sheet in the Prodromus herbarium at Geneva was the basis for that decision. It has two mounted specimens, one labeled A. arenaria Nuttall, collected by Nuttall in “N. Jersey, 1826,” and a second labeled “Atriplex mucronata Rafinesque (A. arenaria Nuttall, N. Jersey) Maritime NY, Rafinesque 1819.” From that information (although the year is 1819, not 1817), and from the description of the taxon, it seems clear that A. arenaria Nuttall is a later synonym of A. mucronata Rafinesque by at least half a year.

H. A. Gleason and A. Cronquist (1991) indicated that this taxon, by whatever name, is “perhaps better treated as a variety of the more tropical Atriplex pentandra (Jacquin) Standley, but the proper nomenclatural combination not yet made.” The present writer agrees with that conclusion, but such subjugation might indicate further contraction into the species of additional closely related taxa, e.g., A. wrightii, which is clearly closely allied as well.

Plants from the coastal states from New England south to New Jersey are much alike and seldom, if ever, display prominent, terminal, naked spikes or panicles with beadlike glomerules of staminate flowers. Plants from Florida westward sometimes have such staminate spikes or panicles.

Specimens from Florida and some from Texas can be distinguished from Atriplex pentandra only with difficulty, especially those individuals with entire leaves. However, the fruiting bracteole length-width ratio and overall shape, with some allowance for overlap, can serve to distinguish most specimens; those of A. mucronata are proportionately longer than broad and, on average, larger.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

At least some early collections were from ballast dumps at harbors on both coasts. It seems probable that the plants were quickly spread inland from initial centers of introduction by birds and more recently along railroads.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4, p. 362. FNA vol. 4, p. 340.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae Chenopodiaceae > Atriplex > subg. Atriplex > sect. Sclerocalymma
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Synonyms A. arenaria, A. cristata var. arenaria, A. pentandra subsp. arenaria
Name authority Rafinesque: Amer. Monthly Mag. & Crit. Rev. 2(2): 119. (1817) Linnaeus: Sp. Pl. ed. 2, 2: 1493. (1763)
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