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quelite

armed saltbush, burscale, tubercled saltbush

Habit Herbs, annual. Shrubs or subshrubs, dioecious, evergreen, mainly 2–10 × 4–10+ dm, woody especially basally, unarmed; branchlets obtusely angled to subterete.
Stems

erect, ascending, or procumbent, much branched, obtusely angled, 1–6 dm, stout, scurfy when young.

Leaves

alternate or proximalmost opposite or subopposite;

petiole to 1.5 cm or sessile;

blade paler abaxially, oblong or oval, lanceolate, or elliptic to broadly obovate or narrowly oblong, 10–40(–60) × (2–)4–20 mm, base rounded to cuneate, margin entire or undulate, rarely with 1–2 teeth, apex rounded to acute, mucronate, thin, often densely white scurfy beneath, grayish green and glabrate above.

persistent, proximal ones opposite, becoming alternate distally, short petiolate or subsessile;

blade oblong to oblong-lanceolate, ovate, obovate, or spatulate, 12–40(–50) × 5–25 mm, base commonly subhastate to cuneate, margin entire or sinuate-dentate to strongly undulate-crisped, apex acute.

Staminate flowers

in interrupted or crowded glomerules 2–4.5 mm thick, in sparsely leafy paniculate spikes to 5+ dm.

Pistillate flowers

fascicled in axils.

few to solitary, in axillary clusters or in crowded or interrupted, often leafy, erect, branched spicate racemes or racemose panicles to 25+ cm.

Seeds

reddish brown, 2 mm.

brown, 1.5–2 mm.

Staminate

glomerules terminal or in dense or interrupted, terminal or axillary, naked spikes or shortly branched panicles.

Fruiting

bracteoles subsessile or with stipes to 1 mm, cuneate-orbiculate, compressed, 4.5–7 × 3.5–5.6 mm, typically longer than wide, united to middle, apex rounded, 3–5-toothed, teeth subequal, sides irregularly tuberculate or with 2 lateral dentate crests, rarely not appendaged.

bracteoles on slender or stout stipes (2–)4–20 mm (or sessile), body broadly elliptic to globose, 6–15 mm and wide, spongy, united to the linear apex, margin deeply laciniate, faces appendaged with flattened to hornlike tubercles to 8 mm.

2n

= 18.

Atriplex mucronata

Atriplex acanthocarpa

Phenology Flowering summer–fall.
Habitat Sandy seashores, salt marshes
Elevation 0 m (0 ft)
Distribution
from FNA
AL; FL; LA; MA; MD; MS; NC; NH; NJ; TX; VA
[BONAP county map]
from FNA
AZ; NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Problems with prior applications of the name Atriplex mucronata lie in the mistaken determination of the place of publication as Amer. Monthly Mag. & Crit. Rev. 2: 176. 1818 (where the name is only mentioned), instead of 2: 119. 1817 (where accompanied by a description and notes). H. M. Hall and F. E. Clements (1923) evidently relied on P. C. Standley’s (1916) interpretation, which indicated the wrong citation (see also A. dioica Rafinesque as an identical example). Hall and Clements applied the name mucronata to their interpretation of A. patula subsp. hastata or to A. hastata (i.e., to A. prostrata according to this treatment).

The treatment of Obione by C. H. B. A. Moquin-Tandon (1849) included “A. mucronata Rafin.!” as a synonym of O. arenaria. Possibly a sheet in the Prodromus herbarium at Geneva was the basis for that decision. It has two mounted specimens, one labeled A. arenaria Nuttall, collected by Nuttall in “N. Jersey, 1826,” and a second labeled “Atriplex mucronata Rafinesque (A. arenaria Nuttall, N. Jersey) Maritime NY, Rafinesque 1819.” From that information (although the year is 1819, not 1817), and from the description of the taxon, it seems clear that A. arenaria Nuttall is a later synonym of A. mucronata Rafinesque by at least half a year.

H. A. Gleason and A. Cronquist (1991) indicated that this taxon, by whatever name, is “perhaps better treated as a variety of the more tropical Atriplex pentandra (Jacquin) Standley, but the proper nomenclatural combination not yet made.” The present writer agrees with that conclusion, but such subjugation might indicate further contraction into the species of additional closely related taxa, e.g., A. wrightii, which is clearly closely allied as well.

Plants from the coastal states from New England south to New Jersey are much alike and seldom, if ever, display prominent, terminal, naked spikes or panicles with beadlike glomerules of staminate flowers. Plants from Florida westward sometimes have such staminate spikes or panicles.

Specimens from Florida and some from Texas can be distinguished from Atriplex pentandra only with difficulty, especially those individuals with entire leaves. However, the fruiting bracteole length-width ratio and overall shape, with some allowance for overlap, can serve to distinguish most specimens; those of A. mucronata are proportionately longer than broad and, on average, larger.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade ovate to rhombic-ovate, elliptic, or spatulate, margin mainly sinuate-dentate; mature fruiting inflorescences ascending to erect; west Texas, s New Mexico, se Arizona
var. acanthocarpa
1. Leaf blade, at least some, hastate-lanceolate, or strongly undulate-crisped; mature fruiting inflorescences mainly horizontally spreading; s Texas, Mexico
var. coahuilensis
Source FNA vol. 4, p. 362. FNA vol. 4, p. 370.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae Chenopodiaceae > Atriplex > subg. Pterochiton
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. acanthocarpa var. acanthocarpa, A. acanthocarpa var. coahuilensis
Synonyms A. arenaria, A. cristata var. arenaria, A. pentandra subsp. arenaria Obione acanthocarpa
Name authority Rafinesque: Amer. Monthly Mag. & Crit. Rev. 2(2): 119. (1817) (Torrey) S. Watson: Proc. Amer. Acad. Arts 9: 117. (1874)
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