Atriplex mucronata
Atriplex joaquiniana
quelite
San Joaquin orach
erect, ascending, or procumbent, much branched, obtusely angled, 1–6 dm, stout, scurfy when young.
alternate or proximalmost opposite or subopposite;
petiole to 1.5 cm or sessile;
blade paler abaxially, oblong or oval, lanceolate, or elliptic to broadly obovate or narrowly oblong, 10–40(–60) × (2–)4–20 mm, base rounded to cuneate, margin entire or undulate, rarely with 1–2 teeth, apex rounded to acute, mucronate, thin, often densely white scurfy beneath, grayish green and glabrate above.
mostly alternate;
petiole 0.5–2.5 cm, becoming shorter or subsessile in distal ones;
blade deltoid to rhombic-ovate or lanceolate, (10–)15–50(–70) × 8–40 mm, base rounded, truncate, or broadly cuneate, margin irregularly sinuate-dentate or repand-dentate to entire, sometimes subhastate, apex obtuse to acute;
distal blades often narrower and sometimes entire except for basal lobes.
in dense or interrupted, naked, simple or paniculate spikes mainly 5–8 mm thick, staminate ones 4-merous.
fascicled in axils.
reddish brown, 2 mm.
dark brown or black, 0.8–1.5 mm;
radicle inferior.
glomerules terminal or in dense or interrupted, terminal or axillary, naked spikes or shortly branched panicles.
bracteoles subsessile or with stipes to 1 mm, cuneate-orbiculate, compressed, 4.5–7 × 3.5–5.6 mm, typically longer than wide, united to middle, apex rounded, 3–5-toothed, teeth subequal, sides irregularly tuberculate or with 2 lateral dentate crests, rarely not appendaged.
bracteoles densely packed on rachis, sessile, ovate-oblong or rounded-deltoid, (2–)2.5–3(–4.5) mm, united only at rounded or truncate base, angled and cristate on faces, thin or spongy-thickened.
= 18.
Atriplex mucronata
Atriplex joaquiniana
Problems with prior applications of the name Atriplex mucronata lie in the mistaken determination of the place of publication as Amer. Monthly Mag. & Crit. Rev. 2: 176. 1818 (where the name is only mentioned), instead of 2: 119. 1817 (where accompanied by a description and notes). H. M. Hall and F. E. Clements (1923) evidently relied on P. C. Standley’s (1916) interpretation, which indicated the wrong citation (see also A. dioica Rafinesque as an identical example). Hall and Clements applied the name mucronata to their interpretation of A. patula subsp. hastata or to A. hastata (i.e., to A. prostrata according to this treatment).
The treatment of Obione by C. H. B. A. Moquin-Tandon (1849) included “A. mucronata Rafin.!” as a synonym of O. arenaria. Possibly a sheet in the Prodromus herbarium at Geneva was the basis for that decision. It has two mounted specimens, one labeled A. arenaria Nuttall, collected by Nuttall in “N. Jersey, 1826,” and a second labeled “Atriplex mucronata Rafinesque (A. arenaria Nuttall, N. Jersey) Maritime NY, Rafinesque 1819.” From that information (although the year is 1819, not 1817), and from the description of the taxon, it seems clear that A. arenaria Nuttall is a later synonym of A. mucronata Rafinesque by at least half a year.
H. A. Gleason and A. Cronquist (1991) indicated that this taxon, by whatever name, is “perhaps better treated as a variety of the more tropical Atriplex pentandra (Jacquin) Standley, but the proper nomenclatural combination not yet made.” The present writer agrees with that conclusion, but such subjugation might indicate further contraction into the species of additional closely related taxa, e.g., A. wrightii, which is clearly closely allied as well.
Plants from the coastal states from New England south to New Jersey are much alike and seldom, if ever, display prominent, terminal, naked spikes or panicles with beadlike glomerules of staminate flowers. Plants from Florida westward sometimes have such staminate spikes or panicles.
Specimens from Florida and some from Texas can be distinguished from Atriplex pentandra only with difficulty, especially those individuals with entire leaves. However, the fruiting bracteole length-width ratio and overall shape, with some allowance for overlap, can serve to distinguish most specimens; those of A. mucronata are proportionately longer than broad and, on average, larger.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Atriplex joaquiniana commonly occurs with Distichlis spicata, Allenrolfea occidentalis, Suaeda moquinii, Frankenia salina, Hordeum depressum, Spergularia macrotheca, and various annual species. H. M. Hall and F. E. Clements (1923) placed the relationship of this very distinctive California endemic with the Atriplex patula complex, with which one can make out a distant affinity. It differs from all other members previously treated within that complex in the small, more or less quadrangular fruiting bracteoles, which are typically crested on one or both elevated faces and have entire or less commonly dentate margins. The bracteoles are borne in very compact simple or paniculate spikes.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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