Atriplex lentiformis
Atriplex californica
big saltbrush, big saltbush, quail bush
California orach, California saltbush
many branched, subterete, 1.5–5 dm, white scurfy when young.
persistent, alternate, petiolate;
blade gray-green, deltate to rhombic, ovate, or oblong-elliptic, 5–50 × 5–50 mm, base truncate to subhastate, margin entire to repand or subhastately lobed, apex rounded to obtuse, scurfy.
numerous, often crowded, alternate or proximalmost opposite;
blade narrowly lanceolate to narrowly oblanceolate or elliptic, 4–20 × 1–5 mm, acute at both ends, gray scurfy.
yellow, in clusters 1–2 mm wide, borne in panicles 0.5–5 dm.
in terminal bracteate spikes, or mixed with pistillate in rather dense axillary clusters, 4-merous.
with less complex panicles.
brown, 0.8–1.6 mm wide.
dark (black), 1–2 mm.
bracteoles sessile, orbiculate to oval, greatly compressed, mainly 3–4.5 mm and wide, crenulate, apex rounded.
bracteoles sessile, rhombic-ovate to ovate, scarcely united, 2.5–4 mm, margin entire, acute.
= 18.
Atriplex lentiformis
Atriplex californica
Materials of big saltbush from the coastal and near coastal regions of California have somewhat broader, merely ovate, rounded leaves, and they have been regarded either at species level as Atriplex breweri S. Watson or at either varietal or subspecific level (see synonymy). The plants intergrade completely in interior situations with typical A. lentiformis, and their recognition at taxonomic level seems superfluous. C. A. Hanson (1962) noted the existence of putative hybrids between A. lentiformis and the herbaceous species A. leucophylla and A. davidsonii. Putative hybrids are also known between this species and A. canescens.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
H. M. Hall and F. E. Clements (1923) placed great emphasis on the inferior radicle, dioecious habit, and free bracts in stating that there are no close relatives of Atriplex californica in North America. P. C. Standley (1916) regarded the radicle as lateral or superior, not inferior, and placed the species at the beginning of his treatment of the American species. Plants of A. californica, however, form a mirror-image, matched pair with A. watsonii, with which they share habit, leaf conformation, staminate glomerules arranged, at least partially, in terminal interrupted spikes, and Kranz anatomy, but from which they differ in the radicle position, valves of the fruiting bracteoles free beyond the middle, monoecious habit, and mostly alternate leaves. The interpretation by Hall and Clements of radicle position as fundamental is made problematic by the apparent random occurrence of a great many morphologic features from place to place within the genus and often within the taxon. Each character fails individually as having definitive taxonomic importance, making difficult or impossible an ultimately satisfactory phylogenetic arrangement. It is not, however, the character that makes the species, rather, it is the entire syndrome of features that constitutes the taxon. Most certainly A. californica is more closely allied to American taxa than to those from other regions of the world.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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