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San Joaquin orach

Garrett's saltbush

Habit Herbs, monoecious or subdioecious, erect, sparsely branched, (1–)3–10 dm; branches obtusely angled, rather rigidly ascending, finely farinose when young. Shrubs or subshrubs, dioecious (rarely monoecious), mainly 2–8(–10) × 5–10+ dm, unarmed.
Leaves

mostly alternate;

petiole 0.5–2.5 cm, becoming shorter or subsessile in distal ones;

blade deltoid to rhombic-ovate or lanceolate, (10–)15–50(–70) × 8–40 mm, base rounded, truncate, or broadly cuneate, margin irregularly sinuate-dentate or repand-dentate to entire, sometimes subhastate, apex obtuse to acute;

distal blades often narrower and sometimes entire except for basal lobes.

opposite or subopposite proximally, or some alternate distally;

petiole 2–6 mm;

blade yellow green, ovate to obovate, elliptic, or orbiculate, 8–55 × 6–32 mm, base obtuse to cuneate, margin entire or repand-dentate, apex rounded to acute, sparingly scurfy.

Flowers

in dense or interrupted, naked, simple or paniculate spikes mainly 5–8 mm thick, staminate ones 4-merous.

Staminate flowers

brown to tan or yellow, in clusters 2–4 mm wide, on panicles 2–10(–15) cm.

Pistillate flowers

in spikes or spicate panicles 4–30 cm.

Seeds

dark brown or black, 0.8–1.5 mm;

radicle inferior.

brown, 2 mm wide.

Fruiting

bracteoles densely packed on rachis, sessile, ovate-oblong or rounded-deltoid, (2–)2.5–3(–4.5) mm, united only at rounded or truncate base, angled and cristate on faces, thin or spongy-thickened.

bracteoles sessile or on stipes to 7 mm, body 4-winged, 6–10 mm and wide, toothed apically, surface smooth, reticulate, or with flattened processes, wings 3–4 mm wide.

2n

= 18.

Atriplex joaquiniana

Atriplex garrettii

Phenology Flowering summer.
Habitat Alkali sink scrub or alkaline grasslands
Elevation 0-200(-300) m (0-700(-1000) ft)
Distribution
from FNA
CA
from FNA
AZ; UT
[BONAP county map]
Discussion

Atriplex joaquiniana commonly occurs with Distichlis spicata, Allenrolfea occidentalis, Suaeda moquinii, Frankenia salina, Hordeum depressum, Spergularia macrotheca, and various annual species. H. M. Hall and F. E. Clements (1923) placed the relationship of this very distinctive California endemic with the Atriplex patula complex, with which one can make out a distant affinity. It differs from all other members previously treated within that complex in the small, more or less quadrangular fruiting bracteoles, which are typically crested on one or both elevated faces and have entire or less commonly dentate margins. The bracteoles are borne in very compact simple or paniculate spikes.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants mainly 2-6 dm; leaves 9 or fewer on first flowering branch of current growth; bract wings denticulate; staminate flowers commonly brown; talus slopes in canyons of the Colorado River, se Utah
var. garrettii
1. Plants mainly 5-10 dm; leaves 7 or more on first flowering branch of current growth; bract wings deeply denticulate; staminate flowers yellow; vicinity of Navajo Bridge and Lee’s Ferry, Arizona
var. navajoensis
Source FNA vol. 4, p. 335. FNA vol. 4, p. 375.
Parent taxa Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis Chenopodiaceae > Atriplex > subg. Pterochiton
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. garrettii var. garrettii, A. garrettii var. navajoensis
Synonyms A. spicata, A. patula subsp. spicata, A. spicata var. lagunita A. canescens subsp. garrettii, A. canescens var. garrettii
Name authority A. Nelson: Proc. Biol. Soc. Wash. 17: 99. (1904) Rydberg: Bull. Torrey Bot. Club 39: 312. (1912)
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