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bract orache, glabrous orach, scotland orache

Bakersfield smallscale, Tulare orach, Tulare saltbush

Habit Herbs, monoecious, prostrate or sprawling, or sometimes erect, branched, (1–)2–10 dm; branches opposite or subopposite. Herbs, erect, simple or sparingly branched, 1.5–4(–10) dm, white scurfy.
Stems

green and striped, often blue-green when fresh, weakly ridged, sparsely scurfy to glabrous.

aging red, terete or obscurely angled, brittle.

Leaves

petiole 0.2–2.5(–3.5) cm;

blade all entire or some or all triangular or lance-hastate with lobes spreading to antrorse, 5–100 × 3–80 mm, base abruptly to narrowly cuneate, entire or irregularly toothed.

alternate or proximalmost opposite;

blade lanceolate to ovate, 6–20 × 4–8 mm, base rounded, margin entire, apex acute to acuminate, gray scurfy.

Flowers

in loose glomerules, arranged in foliose, interrupted spikes or axillary, terminating stems and branches.

Staminate flowers

in small axillary dense glomerules, 4-merous.

Pistillate flowers

solitary or in small axillary clusters, below or mixed with staminate.

Seeds

dimorphic: brown, 2.5–4 mm wide (often the only ones present), or black, (1.2–)1.5–2.9(–3) mm wide;

radicle median, ± antrorse, of brown seed basal and spreading.

dark brown, 1–1.2 mm.

Bracteoles

green, becoming black or reddish to yellow brown, sessile or some short stipitate, venation obscure, ovate-triangular to rhombic-triangular, 5–13 mm, margin united almost to middle, with few irregular teeth or entire, apex abruptly acuminate, faces irregularly muricate, tuberculate, or smooth, inflated, spongy inner layer strongly developed at bracteole base.

Fruiting

bracteoles sessile, rhombic-ovate, acute or acuminate, 2.5–3.5 × 2.2–3 mm, united to middle, thin margin toothed, faces plane, scurfy.

2n

= 18, 36.

Atriplex glabriuscula

Atriplex tularensis

Phenology Flowering summer–fall.
Habitat Alkaline plains, edges of alkali sink
Elevation 90-200 m (300-700 ft)
Distribution
from FNA
CT; MA; ME; NH; PA; AB; MB; NB; NS; PE; QC; Europe
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Members of the Atriplex glabriuscula complex occupy saline or brackish marshes and saline coastal strands mainly in the eastern maritime provinces of Canada, with extensions in similar habitats into the northeastern United States. They are seldom, if ever, ruderal weeds and appear to be indigenous or perhaps early introduced in some part from similar European habitats. The constituent taxa have been regarded at specific level (P. M. Taschereau 1972; I. J. Bassett et al. 1983). They are, however, alike in all major morphologic features, and are apparently closely allied. For those who wish to treat them at specific level, the names are supplied in the synonymy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

H. M. Hall and F. E. Clements (1923) discussed the relationship of this plant to Atriplex cordulata, noting that, “A. tularensis is a much more slender plant, the leaves are narrower in proportion to their length,” and tapering to the base, never at all cordate. They noted further, that “The fruiting bracts are sometimes very similar in shape, but those of tularensis typically end in an abrupt acute tooth not present in cordulata and both bract and seed are always smaller.” Because of its small fruiting bracteoles it was placed within the Pusillae by P. C. Standley (1916). The erect stature, coupled with distinctive spreading-ascending and stiff (at least in dried material) leaves, and foliose bracts apparently distinguish A. tularensis from A. coronata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Bracteoles thin throughout; black seeds 2.0-2.9 mm; bracteoles entire or with tooth on each margin, ovate-triangular
var. acadiensis
1. Bracteoles often if not always spongy-thickened at the base; black seeds rare or lacking, or of various size; bracteoles variously toothed
→ 2
2. Bracteoles sessile or nearly so; inflorescence with leafy bracts throughout; black seeds rare (or lacking), 1.5-2.5 mm; United States and Canada
var. glabriuscula
2. Bracteoles, at least some axillary ones, short stipitate; inflorescence with leafy bracts only near base; black seeds 1.2-2 mm; Canada
var. franktonii
Source FNA vol. 4. FNA vol. 4, p. 359.
Parent taxa Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Pusillae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. glabriuscula var. acadiensis, A. glabriuscula var. franktonii, A. glabriuscula var. glabriuscula
Synonyms A. cordulata var. tularensis, Obione tularensis
Name authority Edmondston: Fl. Shetland, 39. (1845) Coville: Contr. U.S. Natl. Herb. 4: 182, plate 19. (1893)
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