Atriplex glabriuscula
Atriplex suckleyi
bract orache, glabrous orach, scotland orache
Suckley's orach
green and striped, often blue-green when fresh, weakly ridged, sparsely scurfy to glabrous.
terete, often tinged with red, sparsely mealy or glabrous.
petiole 0.2–2.5(–3.5) cm;
blade all entire or some or all triangular or lance-hastate with lobes spreading to antrorse, 5–100 × 3–80 mm, base abruptly to narrowly cuneate, entire or irregularly toothed.
numerous, alternate, sessile;
blade lanceolate to elliptic or less commonly ovate, 7–35 × (2–)4–10(–11) mm, thick and succulent, base acute, margin entire, apex acute or acuminate, glaucous, sparsely farinose when young.
in loose glomerules, arranged in foliose, interrupted spikes or axillary, terminating stems and branches.
in small glomerules, these in distal axils or in short, dense or interrupted, mostly simple, terminal spikes, perianth cup-shaped, frequently pinkish, lobes each with fleshy crest.
solitary or few in leaf axils, calyx within bracteoles of 3 or 4, entire or lobed, distinct, obtuse, hyaline sepals.
dimorphic: brown, 2.5–4 mm wide (often the only ones present), or black, (1.2–)1.5–2.9(–3) mm wide;
radicle median, ± antrorse, of brown seed basal and spreading.
brown, 1.5 × 1.2 mm.
green, becoming black or reddish to yellow brown, sessile or some short stipitate, venation obscure, ovate-triangular to rhombic-triangular, 5–13 mm, margin united almost to middle, with few irregular teeth or entire, apex abruptly acuminate, faces irregularly muricate, tuberculate, or smooth, inflated, spongy inner layer strongly developed at bracteole base.
bracteoles small and difficult to find, sessile, ovate, 2 × 1.5 mm, membranous, united to apex, without appendages, scurfy, each pistil subtended by a perianth.
= 18, 36.
Atriplex glabriuscula
Atriplex suckleyi
Varieties 3 (3 in the flora).
Members of the Atriplex glabriuscula complex occupy saline or brackish marshes and saline coastal strands mainly in the eastern maritime provinces of Canada, with extensions in similar habitats into the northeastern United States. They are seldom, if ever, ruderal weeds and appear to be indigenous or perhaps early introduced in some part from similar European habitats. The constituent taxa have been regarded at specific level (P. M. Taschereau 1972; I. J. Bassett et al. 1983). They are, however, alike in all major morphologic features, and are apparently closely allied. For those who wish to treat them at specific level, the names are supplied in the synonymy.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
At specific rank, the epithet suckleyi clearly has priority. Stutz et al. (1993) resurrected Endolepis Torrey and placed within it two completely disparate species, E. dioica and E. covillei Standley [Atriplex covillei (Standley) J. F. Macbride]. The interpretation of the genus by Stutz et al. stands on the presence of perianth segments subtending the ovary within the fruiting bracteoles, lack of Kranz leaf anatomy, and other more equivocal characteristics. The shared features hardly indicate near affinity, however. The two taxa are otherwise grossly dissimilar. Fundamentally, the genus Endolepis as resurrected by H. C. Stutz et al. stands on the basis of a single character: the presence of perianth segments. Kranz anatomy rises and falls, both within subg. Atriplex and subg. Obione. Thus, coincidence of the non-Kranz criterion is subject to interpretation. Perianth segments subtending the ovary within the enclosing bracteoles, per se, appear to be of independent origin. And, the peculiar nature of the staminate calyx in A. suckleyi (a major determining condition in establishment of the genus Endolepis by Torrey) is not present in A. covillei. Certainly the two species included within Endolepis appear to be as closely allied to other species of Atriplex as they are to each other.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Bracteoles thin throughout; black seeds 2.0-2.9 mm; bracteoles entire or with tooth on each margin, ovate-triangular | var. acadiensis |
1. Bracteoles often if not always spongy-thickened at the base; black seeds rare or lacking, or of various size; bracteoles variously toothed | → 2 |
2. Bracteoles sessile or nearly so; inflorescence with leafy bracts throughout; black seeds rare (or lacking), 1.5-2.5 mm; United States and Canada | var. glabriuscula |
2. Bracteoles, at least some axillary ones, short stipitate; inflorescence with leafy bracts only near base; black seeds 1.2-2 mm; Canada | var. franktonii |
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