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bract orache, glabrous orach, scotland orache

sack saltbush, sack saltweed, stalk orach, stalk orache

Habit Herbs, monoecious, prostrate or sprawling, or sometimes erect, branched, (1–)2–10 dm; branches opposite or subopposite. Herbs, erect, forming rounded clumps.
Stems

green and striped, often blue-green when fresh, weakly ridged, sparsely scurfy to glabrous.

mainly 0.5–4(–5) dm;

stems usually branched from base, terete, stout, angled;

herbage scurfy.

Leaves

petiole 0.2–2.5(–3.5) cm;

blade all entire or some or all triangular or lance-hastate with lobes spreading to antrorse, 5–100 × 3–80 mm, base abruptly to narrowly cuneate, entire or irregularly toothed.

alternate or proximalmost subopposite, short petiolate or distal ones sessile;

petiole 2–5 mm;

blade cordate-ovate or subreniform to ovate to deltoid-ovate or oval, mainly 6–40 × 4–30 mm, base truncate to subcordate or broadly cuneate, margin entire or in some subhastately lobed or sometimes undulate-dentate, apex acute to rounded.

Flowers

in loose glomerules, arranged in foliose, interrupted spikes or axillary, terminating stems and branches.

Pistillate flowers

usually in fascicles of 1–3 in proximal axils.

Seeds

dimorphic: brown, 2.5–4 mm wide (often the only ones present), or black, (1.2–)1.5–2.9(–3) mm wide;

radicle median, ± antrorse, of brown seed basal and spreading.

brownish to whitish, 1.5–2.3 mm.

Bracteoles

green, becoming black or reddish to yellow brown, sessile or some short stipitate, venation obscure, ovate-triangular to rhombic-triangular, 5–13 mm, margin united almost to middle, with few irregular teeth or entire, apex abruptly acuminate, faces irregularly muricate, tuberculate, or smooth, inflated, spongy inner layer strongly developed at bracteole base.

Staminate

glomerules in distal axils or in short, naked, terminal (early deciduous) panicles;

flowers with 5-parted calyx.

Fruiting

bracteoles monomorphic or dimorphic, larger ones on stipes (2–)4–8(–15) mm, others sessile, united at base, round-triangular or suborbicular, 4–6 mm, irregularly and coarsely dentate and with ± densely beset with flat, cristate, or hornlike appendages, smaller bracteoles (lacking or rare in var. asterocarpa) in same axils, oblong to cuneate, 3–4 mm, apex truncate, dentate only at summit, faces smooth.

2n

= 18, 36.

= 18.

Atriplex glabriuscula

Atriplex saccaria

Distribution
from FNA
CT; MA; ME; NH; PA; AB; MB; NB; NS; PE; QC; Europe
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CO; NM; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Members of the Atriplex glabriuscula complex occupy saline or brackish marshes and saline coastal strands mainly in the eastern maritime provinces of Canada, with extensions in similar habitats into the northeastern United States. They are seldom, if ever, ruderal weeds and appear to be indigenous or perhaps early introduced in some part from similar European habitats. The constituent taxa have been regarded at specific level (P. M. Taschereau 1972; I. J. Bassett et al. 1983). They are, however, alike in all major morphologic features, and are apparently closely allied. For those who wish to treat them at specific level, the names are supplied in the synonymy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Bracteoles thin throughout; black seeds 2.0-2.9 mm; bracteoles entire or with tooth on each margin, ovate-triangular
var. acadiensis
1. Bracteoles often if not always spongy-thickened at the base; black seeds rare or lacking, or of various size; bracteoles variously toothed
→ 2
2. Bracteoles sessile or nearly so; inflorescence with leafy bracts throughout; black seeds rare (or lacking), 1.5-2.5 mm; United States and Canada
var. glabriuscula
2. Bracteoles, at least some axillary ones, short stipitate; inflorescence with leafy bracts only near base; black seeds 1.2-2 mm; Canada
var. franktonii
1. Leaves mainly ovate to cordate in profile, the distalmost bracteate ones (and sometimes all) often broadly so in age; fruiting bracteoles with stipes to 10 mm, but seldom exceeding 6 mm; Arizona, New Mexico, Wyoming, s to Texas
var. saccaria
1. Leaves oval to rhombic, the bases truncate to cuneate-attenuate; fruiting bracteoles with stipes of various length; sw Utah and adjacent states
→ 2
2. Leaves oval to suborbiculate or less commonly elliptic; fruiting bracteoles with hornlike appendages, borne on stipes mainly 2-6 mm
var. asterocarpa
2. Leaves rhombic to oval or triangular, base cuneate-attenuate; fruiting bracteoles with flattened processes, borne on stipes mainly 2-15 mm (sessile in some)
var. cornuta
Source FNA vol. 4. FNA vol. 4.
Parent taxa Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Saccariae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. glabriuscula var. acadiensis, A. glabriuscula var. franktonii, A. glabriuscula var. glabriuscula
A. saccaria var. asterocarpa, A. saccaria var. cornuta, A. saccaria var. saccaria
Synonyms A. truncata var. saccaria, Obione saccaria
Name authority Edmondston: Fl. Shetland, 39. (1845) S. Watson: Proc. Amer. Acad. Arts 9: 112. (1874)
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