Atriplex glabriuscula |
Atriplex pentandra |
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bract orache, glabrous orach, scotland orache |
seas hore orach |
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Habit | Herbs, monoecious, prostrate or sprawling, or sometimes erect, branched, (1–)2–10 dm; branches opposite or subopposite. | Herbs, annual or perennial, sprawling to erect, often suffrutescent at the base, much branched and clump-forming, 3–10 dm. | ||||||||
Stems | green and striped, often blue-green when fresh, weakly ridged, sparsely scurfy to glabrous. |
obtusely angled, finely scurfy when young. |
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Leaves | petiole 0.2–2.5(–3.5) cm; blade all entire or some or all triangular or lance-hastate with lobes spreading to antrorse, 5–100 × 3–80 mm, base abruptly to narrowly cuneate, entire or irregularly toothed. |
sessile or short petiolate; blade paler abaxially, oblong or rhombic-ovate to broadly obovate or narrowly oblong or narrowly elliptic, 10–30 × (1–)3–15 mm, thin, base rounded to cuneate, margin repand-dentate or sinuate-dentate to undulate or distal ones or all of them entire, apex rounded to acute, mucronate, densely white scurfy abaxially, grayish green and usually glabrate adaxially. |
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Flowers | in loose glomerules, arranged in foliose, interrupted spikes or axillary, terminating stems and branches. |
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Staminate flowers | in short, dense, naked terminal spikes or panicles; calyx 5-cleft, lobes green keeled. |
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Pistillate flowers | fascicled in axils. |
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Seeds | dimorphic: brown, 2.5–4 mm wide (often the only ones present), or black, (1.2–)1.5–2.9(–3) mm wide; radicle median, ± antrorse, of brown seed basal and spreading. |
brown, 1–1.5 mm. |
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Bracteoles | green, becoming black or reddish to yellow brown, sessile or some short stipitate, venation obscure, ovate-triangular to rhombic-triangular, 5–13 mm, margin united almost to middle, with few irregular teeth or entire, apex abruptly acuminate, faces irregularly muricate, tuberculate, or smooth, inflated, spongy inner layer strongly developed at bracteole base. |
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Fruiting | bracteoles sessile or with stipes to 0.5 mm, broadly cuneate-orbiculate, compressed, (2–) 2.5–4.5 × (1.5–)2.6–5 mm, usually as broad as or broader than long, much thickened at maturity, united only at truncate or broadly cuneate base, margin deeply and acutely dentate, faces with 2, sometimes swollen, dentate crests or covered with irregular, conic-acute, corky tubercles, seldom smooth. |
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2n | = 18, 36. |
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Atriplex glabriuscula |
Atriplex pentandra |
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Phenology | Flowering summer–fall. | |||||||||
Habitat | Sandy seashores, coastal salt marshes | |||||||||
Elevation | 0-50 m (0-200 ft) | |||||||||
Distribution |
CT; MA; ME; NH; PA; AB; MB; NB; NS; PE; QC; Europe
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AL; CT; FL; GA; LA; MA; MS; NC; SC; TX; West Indies; South America (Venezuela and Colombia to Peru) |
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Discussion | Varieties 3 (3 in the flora). Members of the Atriplex glabriuscula complex occupy saline or brackish marshes and saline coastal strands mainly in the eastern maritime provinces of Canada, with extensions in similar habitats into the northeastern United States. They are seldom, if ever, ruderal weeds and appear to be indigenous or perhaps early introduced in some part from similar European habitats. The constituent taxa have been regarded at specific level (P. M. Taschereau 1972; I. J. Bassett et al. 1983). They are, however, alike in all major morphologic features, and are apparently closely allied. For those who wish to treat them at specific level, the names are supplied in the synonymy. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
D. S. Correll and M. C. Johnston (1970), treated both Atriplex texana and A. wardii at the specific level. The diagnostic features used to distinguish them, dentate versus entire leaf blades and smooth versus tuberculate faces of fruiting bracteoles, fail singly and in combination. Both taxa were regarded by H. M. Hall and F. E. Clements (1923) as variants of typical A. pentandra, and this worker tentatively agrees with those authors’ conclusions. Specimens from Galveston, Texas—the type locality of A. wardii—are apparently intermediate with A. mucronata (see below) and form the basis on which A. wardii was founded. Some of those specimens have been annotated by me and others as A. mucronata. The plants have entire leaves more closely matching those of A. mucronata, but the small fruiting bracteoles, though usually lacking tubercles on the faces, are of similar proportions to those of typical A. pentandra. Atriplex pentandra and A. mucronata are certainly very closely allied. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4. | FNA vol. 4, p. 362. | ||||||||
Parent taxa | Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis | Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Axyris pentandra, A. texana, A. wardii | |||||||||
Name authority | Edmondston: Fl. Shetland, 39. (1845) | (Jacquin) Standley: in N. L. Britton et al., N. Amer. Fl. 21: 54. (1916) | ||||||||
Web links |