Atriplex glabriuscula
Atriplex mucronata
bract orache, glabrous orach, scotland orache
quelite
green and striped, often blue-green when fresh, weakly ridged, sparsely scurfy to glabrous.
erect, ascending, or procumbent, much branched, obtusely angled, 1–6 dm, stout, scurfy when young.
petiole 0.2–2.5(–3.5) cm;
blade all entire or some or all triangular or lance-hastate with lobes spreading to antrorse, 5–100 × 3–80 mm, base abruptly to narrowly cuneate, entire or irregularly toothed.
alternate or proximalmost opposite or subopposite;
petiole to 1.5 cm or sessile;
blade paler abaxially, oblong or oval, lanceolate, or elliptic to broadly obovate or narrowly oblong, 10–40(–60) × (2–)4–20 mm, base rounded to cuneate, margin entire or undulate, rarely with 1–2 teeth, apex rounded to acute, mucronate, thin, often densely white scurfy beneath, grayish green and glabrate above.
in loose glomerules, arranged in foliose, interrupted spikes or axillary, terminating stems and branches.
fascicled in axils.
dimorphic: brown, 2.5–4 mm wide (often the only ones present), or black, (1.2–)1.5–2.9(–3) mm wide;
radicle median, ± antrorse, of brown seed basal and spreading.
reddish brown, 2 mm.
green, becoming black or reddish to yellow brown, sessile or some short stipitate, venation obscure, ovate-triangular to rhombic-triangular, 5–13 mm, margin united almost to middle, with few irregular teeth or entire, apex abruptly acuminate, faces irregularly muricate, tuberculate, or smooth, inflated, spongy inner layer strongly developed at bracteole base.
glomerules terminal or in dense or interrupted, terminal or axillary, naked spikes or shortly branched panicles.
bracteoles subsessile or with stipes to 1 mm, cuneate-orbiculate, compressed, 4.5–7 × 3.5–5.6 mm, typically longer than wide, united to middle, apex rounded, 3–5-toothed, teeth subequal, sides irregularly tuberculate or with 2 lateral dentate crests, rarely not appendaged.
= 18, 36.
= 18.
Atriplex glabriuscula
Atriplex mucronata
Varieties 3 (3 in the flora).
Members of the Atriplex glabriuscula complex occupy saline or brackish marshes and saline coastal strands mainly in the eastern maritime provinces of Canada, with extensions in similar habitats into the northeastern United States. They are seldom, if ever, ruderal weeds and appear to be indigenous or perhaps early introduced in some part from similar European habitats. The constituent taxa have been regarded at specific level (P. M. Taschereau 1972; I. J. Bassett et al. 1983). They are, however, alike in all major morphologic features, and are apparently closely allied. For those who wish to treat them at specific level, the names are supplied in the synonymy.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Problems with prior applications of the name Atriplex mucronata lie in the mistaken determination of the place of publication as Amer. Monthly Mag. & Crit. Rev. 2: 176. 1818 (where the name is only mentioned), instead of 2: 119. 1817 (where accompanied by a description and notes). H. M. Hall and F. E. Clements (1923) evidently relied on P. C. Standley’s (1916) interpretation, which indicated the wrong citation (see also A. dioica Rafinesque as an identical example). Hall and Clements applied the name mucronata to their interpretation of A. patula subsp. hastata or to A. hastata (i.e., to A. prostrata according to this treatment).
The treatment of Obione by C. H. B. A. Moquin-Tandon (1849) included “A. mucronata Rafin.!” as a synonym of O. arenaria. Possibly a sheet in the Prodromus herbarium at Geneva was the basis for that decision. It has two mounted specimens, one labeled A. arenaria Nuttall, collected by Nuttall in “N. Jersey, 1826,” and a second labeled “Atriplex mucronata Rafinesque (A. arenaria Nuttall, N. Jersey) Maritime NY, Rafinesque 1819.” From that information (although the year is 1819, not 1817), and from the description of the taxon, it seems clear that A. arenaria Nuttall is a later synonym of A. mucronata Rafinesque by at least half a year.
H. A. Gleason and A. Cronquist (1991) indicated that this taxon, by whatever name, is “perhaps better treated as a variety of the more tropical Atriplex pentandra (Jacquin) Standley, but the proper nomenclatural combination not yet made.” The present writer agrees with that conclusion, but such subjugation might indicate further contraction into the species of additional closely related taxa, e.g., A. wrightii, which is clearly closely allied as well.
Plants from the coastal states from New England south to New Jersey are much alike and seldom, if ever, display prominent, terminal, naked spikes or panicles with beadlike glomerules of staminate flowers. Plants from Florida westward sometimes have such staminate spikes or panicles.
Specimens from Florida and some from Texas can be distinguished from Atriplex pentandra only with difficulty, especially those individuals with entire leaves. However, the fruiting bracteole length-width ratio and overall shape, with some allowance for overlap, can serve to distinguish most specimens; those of A. mucronata are proportionately longer than broad and, on average, larger.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Bracteoles thin throughout; black seeds 2.0-2.9 mm; bracteoles entire or with tooth on each margin, ovate-triangular | var. acadiensis |
1. Bracteoles often if not always spongy-thickened at the base; black seeds rare or lacking, or of various size; bracteoles variously toothed | → 2 |
2. Bracteoles sessile or nearly so; inflorescence with leafy bracts throughout; black seeds rare (or lacking), 1.5-2.5 mm; United States and Canada | var. glabriuscula |
2. Bracteoles, at least some axillary ones, short stipitate; inflorescence with leafy bracts only near base; black seeds 1.2-2 mm; Canada | var. franktonii |
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