Atriplex glabriuscula |
Atriplex joaquiniana |
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bract orache, glabrous orach, scotland orache |
San Joaquin orach |
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Habit | Herbs, monoecious, prostrate or sprawling, or sometimes erect, branched, (1–)2–10 dm; branches opposite or subopposite. | Herbs, monoecious or subdioecious, erect, sparsely branched, (1–)3–10 dm; branches obtusely angled, rather rigidly ascending, finely farinose when young. | ||||||||
Stems | green and striped, often blue-green when fresh, weakly ridged, sparsely scurfy to glabrous. |
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Leaves | petiole 0.2–2.5(–3.5) cm; blade all entire or some or all triangular or lance-hastate with lobes spreading to antrorse, 5–100 × 3–80 mm, base abruptly to narrowly cuneate, entire or irregularly toothed. |
mostly alternate; petiole 0.5–2.5 cm, becoming shorter or subsessile in distal ones; blade deltoid to rhombic-ovate or lanceolate, (10–)15–50(–70) × 8–40 mm, base rounded, truncate, or broadly cuneate, margin irregularly sinuate-dentate or repand-dentate to entire, sometimes subhastate, apex obtuse to acute; distal blades often narrower and sometimes entire except for basal lobes. |
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Flowers | in loose glomerules, arranged in foliose, interrupted spikes or axillary, terminating stems and branches. |
in dense or interrupted, naked, simple or paniculate spikes mainly 5–8 mm thick, staminate ones 4-merous. |
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Seeds | dimorphic: brown, 2.5–4 mm wide (often the only ones present), or black, (1.2–)1.5–2.9(–3) mm wide; radicle median, ± antrorse, of brown seed basal and spreading. |
dark brown or black, 0.8–1.5 mm; radicle inferior. |
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Bracteoles | green, becoming black or reddish to yellow brown, sessile or some short stipitate, venation obscure, ovate-triangular to rhombic-triangular, 5–13 mm, margin united almost to middle, with few irregular teeth or entire, apex abruptly acuminate, faces irregularly muricate, tuberculate, or smooth, inflated, spongy inner layer strongly developed at bracteole base. |
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Fruiting | bracteoles densely packed on rachis, sessile, ovate-oblong or rounded-deltoid, (2–)2.5–3(–4.5) mm, united only at rounded or truncate base, angled and cristate on faces, thin or spongy-thickened. |
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2n | = 18, 36. |
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Atriplex glabriuscula |
Atriplex joaquiniana |
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Phenology | Flowering summer. | |||||||||
Habitat | Alkali sink scrub or alkaline grasslands | |||||||||
Elevation | 0-200(-300) m (0-700(-1000) ft) | |||||||||
Distribution |
CT; MA; ME; NH; PA; AB; MB; NB; NS; PE; QC; Europe
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CA |
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Discussion | Varieties 3 (3 in the flora). Members of the Atriplex glabriuscula complex occupy saline or brackish marshes and saline coastal strands mainly in the eastern maritime provinces of Canada, with extensions in similar habitats into the northeastern United States. They are seldom, if ever, ruderal weeds and appear to be indigenous or perhaps early introduced in some part from similar European habitats. The constituent taxa have been regarded at specific level (P. M. Taschereau 1972; I. J. Bassett et al. 1983). They are, however, alike in all major morphologic features, and are apparently closely allied. For those who wish to treat them at specific level, the names are supplied in the synonymy. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Atriplex joaquiniana commonly occurs with Distichlis spicata, Allenrolfea occidentalis, Suaeda moquinii, Frankenia salina, Hordeum depressum, Spergularia macrotheca, and various annual species. H. M. Hall and F. E. Clements (1923) placed the relationship of this very distinctive California endemic with the Atriplex patula complex, with which one can make out a distant affinity. It differs from all other members previously treated within that complex in the small, more or less quadrangular fruiting bracteoles, which are typically crested on one or both elevated faces and have entire or less commonly dentate margins. The bracteoles are borne in very compact simple or paniculate spikes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4. | FNA vol. 4, p. 335. | ||||||||
Parent taxa | Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis | Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | A. spicata, A. patula subsp. spicata, A. spicata var. lagunita | |||||||||
Name authority | Edmondston: Fl. Shetland, 39. (1845) | A. Nelson: Proc. Biol. Soc. Wash. 17: 99. (1904) | ||||||||
Web links |