Atriplex glabriuscula |
Atriplex canescens |
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bract orache, glabrous orach, scotland orache |
four-wing saltbu sh, four-wing shad-scale, hoary saltbush, shadscale, wingscale |
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Habit | Herbs, monoecious, prostrate or sprawling, or sometimes erect, branched, (1–)2–10 dm; branches opposite or subopposite. | Shrubs, dioecious or rarely monoecious, mainly 8–20 dm, as wide or wider, not especially armed. | ||||||||||||||||||||
Stems | green and striped, often blue-green when fresh, weakly ridged, sparsely scurfy to glabrous. |
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Leaves | petiole 0.2–2.5(–3.5) cm; blade all entire or some or all triangular or lance-hastate with lobes spreading to antrorse, 5–100 × 3–80 mm, base abruptly to narrowly cuneate, entire or irregularly toothed. |
persistent, alternate, sessile or nearly so, blade linear to oblanceolate, oblong, or obovate, mainly 10–40 × 3–8 mm, margin entire, apex retuse to obtuse. |
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Flowers | in loose glomerules, arranged in foliose, interrupted spikes or axillary, terminating stems and branches. |
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Staminate flowers | yellow (rarely brown), in clusters 2–3 mm wide, borne in panicles 3–15 cm. |
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Pistillate flowers | borne in panicles 5–40 cm. |
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Seeds | dimorphic: brown, 2.5–4 mm wide (often the only ones present), or black, (1.2–)1.5–2.9(–3) mm wide; radicle median, ± antrorse, of brown seed basal and spreading. |
1.5–2.5 mm wide. |
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Bracteoles | green, becoming black or reddish to yellow brown, sessile or some short stipitate, venation obscure, ovate-triangular to rhombic-triangular, 5–13 mm, margin united almost to middle, with few irregular teeth or entire, apex abruptly acuminate, faces irregularly muricate, tuberculate, or smooth, inflated, spongy inner layer strongly developed at bracteole base. |
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Fruiting | bracteoles 8–25 mm, as wide, on stipes 1–8 mm, with 4 prominent wings extending the bract length, united throughout, wings dentate to entire, apex toothed, surface of wings and body smooth or reticulate. |
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2n | = 18, 36. |
= 18, 36+. |
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Atriplex glabriuscula |
Atriplex canescens |
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Distribution |
CT; MA; ME; NH; PA; AB; MB; NB; NS; PE; QC; Europe
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AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; Mexico
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Discussion | Varieties 3 (3 in the flora). Members of the Atriplex glabriuscula complex occupy saline or brackish marshes and saline coastal strands mainly in the eastern maritime provinces of Canada, with extensions in similar habitats into the northeastern United States. They are seldom, if ever, ruderal weeds and appear to be indigenous or perhaps early introduced in some part from similar European habitats. The constituent taxa have been regarded at specific level (P. M. Taschereau 1972; I. J. Bassett et al. 1983). They are, however, alike in all major morphologic features, and are apparently closely allied. For those who wish to treat them at specific level, the names are supplied in the synonymy. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
This species forms hybrids with Atriplex confertifolia and A. gardneri varieties (see var. bonnevillensis). Materials from the vicinity of the type locality of the species in South Dakota are low subherbaceous plants that differ from our shrubby tall material. However, the type area is presently covered with water from a dam on the Missouri River, and it is not possible to exclude the possibility of A. canescens as it has been interpreted for the past century to have existed at that site during the Lewis and Clark Expedition, if that is indeed where the lectotype was collected. Varieties 4 (4 in the flora). Nuttall’s new combination Atriplex canescens was based on Calligonum canescens Pursh. Watson based his new name A. nuttallii directly on A. canescens Nuttall, i.e., including the citation of Nuttall’s “Genera, 1. 197,” and on its basionym, C. canescens. The name A. nuttallii is thus a nomenclatural synonym of A. canescens and was thus illegitimate at its inception. It cannot be resurrected by even the most sophisticated arguments. A sheet of Atriplex canescens, noted as “a shrub,” taken by Nuttall on the 1810 Missouri River expedition is extant in the Lambert herbarium (PH). It bears several, obviously shrubby, staminate flowering branches, but the only pistillate branch is very immature. The name A. gardneri, also cited provisionally by Watson within the concept of A. nuttallii, clearly has priority over other names for that widely distributed species complex. Attempts at leptotypification of the name nuttallii by J. McNeill et al. (1983) and H. C. Stutz and S. C. Sanderson (1998) are both superfluous, the name being illegitimate. C. A. Hanson (1962) noted the similarity between the occasional wingless fruiting bracteoles of Atriplex canescens and A. gardneri var. falcata. He noted further that the bracts of both species lack lateral teeth subtending the terminal ones, have terminal teeth united half their length, and have indurate bracts. Whether such similarity indicates relationship or mere coincidence is open to question. However, A. canescens is known to form hybrids with most, if not all, portions of the gardneri complex and with other woody species whose range it overlaps as well. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4. | FNA vol. 4. | ||||||||||||||||||||
Parent taxa | Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis | Chenopodiaceae > Atriplex > subg. Pterochiton | ||||||||||||||||||||
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Synonyms | Calligonum canescens, A. nuttallii | |||||||||||||||||||||
Name authority | Edmondston: Fl. Shetland, 39. (1845) | (Pursh) Nuttall: Gen. N. Amer. Pl. 1: 197. (1818) | ||||||||||||||||||||
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