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bract orache, glabrous orach, scotland orache

maidenhair spleenwort, silver orach, silver orache, silver saltbush, silverscale, silverscale orache, silverscale saltbush, silvery orache

Habit Herbs, monoecious, prostrate or sprawling, or sometimes erect, branched, (1–)2–10 dm; branches opposite or subopposite. Herbs, simple or freely branched, 0.5–6 dm; branches rather stout, angled, scurfy when young.
Stems

green and striped, often blue-green when fresh, weakly ridged, sparsely scurfy to glabrous.

Leaves

petiole 0.2–2.5(–3.5) cm;

blade all entire or some or all triangular or lance-hastate with lobes spreading to antrorse, 5–100 × 3–80 mm, base abruptly to narrowly cuneate, entire or irregularly toothed.

often opposite proximally, petiolate or distal bracteate ones subsessile, blade lance-ovate, lanceolate, deltoid, or cordate, 5–75 × 4–50(–75) mm, base subhastate or obtuse to acute, margin entire or essentially so, sometimes closely repand-dentate, apex obtuse to acute or rounded, scurfy (glabrous).

Flowers

in loose glomerules, arranged in foliose, interrupted spikes or axillary, terminating stems and branches.

in axillary glomerules and terminal, interrupted spikes.

Staminate flowers

borne in distal axils, or in short dense spikes or panicles, or intermixed with pistillate, with 4–5-parted calyx.

Seeds

dimorphic: brown, 2.5–4 mm wide (often the only ones present), or black, (1.2–)1.5–2.9(–3) mm wide;

radicle median, ± antrorse, of brown seed basal and spreading.

brown, 1.5–2 mm wide;

radicle superior or lateral.

Bracteoles

green, becoming black or reddish to yellow brown, sessile or some short stipitate, venation obscure, ovate-triangular to rhombic-triangular, 5–13 mm, margin united almost to middle, with few irregular teeth or entire, apex abruptly acuminate, faces irregularly muricate, tuberculate, or smooth, inflated, spongy inner layer strongly developed at bracteole base.

Fruiting

bracteoles sessile, subsessile, or stipitate (stipe 0.5–5 mm), cuneate-orbicular, (2.5–)4–11.2 × 2–8.8(–14) mm, margin foliaceous below apex, subentire or dentate to laciniate, face smooth, tuberculate, or crested, processes sometimes again toothed, teeth then aligned with axis of process.

2n

= 18, 36.

= 18, 36, 54.

Atriplex glabriuscula

Atriplex argentea

Distribution
from FNA
CT; MA; ME; NH; PA; AB; MB; NB; NS; PE; QC; Europe
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB; SK; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Members of the Atriplex glabriuscula complex occupy saline or brackish marshes and saline coastal strands mainly in the eastern maritime provinces of Canada, with extensions in similar habitats into the northeastern United States. They are seldom, if ever, ruderal weeds and appear to be indigenous or perhaps early introduced in some part from similar European habitats. The constituent taxa have been regarded at specific level (P. M. Taschereau 1972; I. J. Bassett et al. 1983). They are, however, alike in all major morphologic features, and are apparently closely allied. For those who wish to treat them at specific level, the names are supplied in the synonymy.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 5 (5 in the flora).

Herbarium materials have tended to represent a catchall for annual specimens not readily assignable to other taxa. Indeed, the distinguishing features of the Atriplex argentea complex are shared singly and often in combination with other taxa. Only by use of combinations of features can this taxon be defined. Those features, with much variation, center around the broad, typically ovate to deltoid leaf blades (often definitely 3-veined) and more-or-less compressed, sessile to subsessile (or short stipitate), fruiting bracteoles on which the marginal processes, or teeth, are mainly aligned with the plane compression, and with the faces quite smooth to variously appendaged. Still some specimens are apparently intermediate with other species, especially with the closely allied A. saccaria, with which it is at least partially sympatric.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Bracteoles thin throughout; black seeds 2.0-2.9 mm; bracteoles entire or with tooth on each margin, ovate-triangular
var. acadiensis
1. Bracteoles often if not always spongy-thickened at the base; black seeds rare or lacking, or of various size; bracteoles variously toothed
→ 2
2. Bracteoles sessile or nearly so; inflorescence with leafy bracts throughout; black seeds rare (or lacking), 1.5-2.5 mm; United States and Canada
var. glabriuscula
2. Bracteoles, at least some axillary ones, short stipitate; inflorescence with leafy bracts only near base; black seeds 1.2-2 mm; Canada
var. franktonii
1. Plants low, rounded, mainly 1-3(-4.5) dm; fruiting bracteoles sessile or rarely some short stipitate, the appendages mainly acuminate or acute-attenuate; Nevada and adjacent se California, se Oregon
→ 2
1. Plants rather strict and often taller, mainly 2-4(-5) dm; fruiting bracteoles, at least some, short stipitate; appendages acute to less commonly acuminate to attenuate; other distribution
→ 3
2. Leaf blades typically, but not always, constricted to the petiole; herbage merely scurfy, or with elongate trichomes mainly on the fruiting bracteoles; Nevada (except the southern part), and adjacent se California and se Oregon
var. hillmanii
2. Leaf blades typically, if not always, cuneate to a winged petiole or merely sessile base; herbage copiously covered with elongate trichomes throughout, or mostly on the stems and fruiting bracteoles; Nye County, Nevada and adjacent Inyo County, California
var. longitrichoma
3. Leaf blades elliptic to oval, attenuate to a cuneate base; sw Colorado, se Utah, and nw New Mexico
var. rydbergii
3. Leaf blades triangular-ovate to oval, base broadly obtuse to acute or less commonly cuneate; various or other distribution
→ 4
4. Distal leaves short petiolate; proximalmost leaves alternate; plants mostly less than 4 dm; British Columbia to Manitoba, s through Montana, North Dakota, South Dakota, Wyoming, Nebraska, Idaho, Utah, e Nevada, Colorado, w Kansas, ne Arizona, nw New Mexico
var. argentea
4. Distal leaves sessile; proximalmost leaves opposite; plants mostly over 4 dm; California, s Nevada, Arizona, New Mexico, w Oklahoma, w and n Texas
var. mohavensis
Source FNA vol. 4. FNA vol. 4.
Parent taxa Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Argenteae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. glabriuscula var. acadiensis, A. glabriuscula var. franktonii, A. glabriuscula var. glabriuscula
A. argentea var. argentea, A. argentea var. hillmanii, A. argentea var. longitrichoma, A. argentea var. mohavensis, A. argentea var. rydbergii
Synonyms Obione argentea
Name authority Edmondston: Fl. Shetland, 39. (1845) Nuttall: Gen. N. Amer. Pl. 1: 198. (1818)
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