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wheelscale, wheelscale orach, wheelscale saltbush, white-scale saltbush

spinescale saltbush, spiny saltbush

Habit Herbs, annual (or sometimes perennial?). Shrubs, dioecious, erect, intricately much branched, mainly 3–15 dm; branchlets terete, becoming rigid and spinose.
Stems

ascending or procumbent to erect, stramineous or whitish, simple or much branched at base, obtusely angled in age, mainly 0.5–4.5 dm, slender or stout, scurfy to glabrate.

Leaves

many, subsessile or shortly petiolate;

blade elliptic to spatulate, oblanceolate, oblong, or obovate, 5–30(–35) × 2–8(–12) mm, base cuneate to attenuate, margin entire or irregularly dentate, densely scurfy abaxially, usually green and glabrate adaxially.

short petiolate to sessile;

blade ovate-deltate to elliptic or spatulate, (5–)10–27 mm, entire or subhastate.

Staminate flowers

with 3–5-parted perianth.

borne in small axillary glomerules.

Pistillate flowers

intermixed with staminate in small axillary clusters.

borne solitary or few in bract axils on short, spinose, lateral branchlets of a paniculate inflorescence.

Seeds

brown, 1–1.5 mm wide.

reddish brown, 2–2.8 mm.

Fruiting

bracteoles subsessile or short stipitate, orbiculate, strongly compressed, 2–4 mm and as wide, united except at thin margin, margin dentate, terminal teeth often prominent, faces smooth or with cristate appendages (thornberi phase).

bracteoles sessile or nearly so, body globose, connate, constricted below oblong to orbicular wings, 7–15 ×3.5–10 mm, entire or obscurely dentate, faces smooth to sparingly cristate.

Atriplex elegans

Atriplex spinifera

Phenology Flowering summer–fall.
Habitat Xeric saline substrates, with mixed salt desert shrubs
Elevation 30-1300 m (100-4300 ft)
Distribution
from FNA
AZ; CA; NM; NV; TX; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Spiny saltbush apparently forms occasional hybrids with phases of Atriplex canescens, as indicated by the presence of wings on some of the fruiting bracteoles.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruiting bracteole margin dentate to incised, teeth 0.5-1 mm; bracteole not especially samaralike; s California e to Texas, and Mexico (Chihuahua, Sonora)
var. elegans
1. Fruiting bracteole margin finely toothed, teeth 0.3-0.5 mm; bracteole appearing samaralike; s Cali- fornia and w Arizona
var. fasciculata
Source FNA vol. 4, p. 364. FNA vol. 4, p. 377.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae Chenopodiaceae > Atriplex > subg. Pterochiton
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. elegans var. elegans, A. elegans var. fasciculata
Synonyms Obione elegans
Name authority (Moquin-Tandon) D. Dietrich: Syn. Pl. 5: 537. (1852) J. F. Macbride: Contr. Gray Herb. 53: 11. (1918)
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