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wheelscale, wheelscale orach, wheelscale saltbush, white-scale saltbush

Coulter's orach, Coulter's orache, Coulter's saltbush

Habit Herbs, annual (or sometimes perennial?). Herbs, perennial, sometimes flowering as an annual, spreading 0.7–10 dm, slightly woody at base.
Stems

ascending or procumbent to erect, stramineous or whitish, simple or much branched at base, obtusely angled in age, mainly 0.5–4.5 dm, slender or stout, scurfy to glabrate.

frequently tinged with red, much branched, sparsely scurfy.

Leaves

many, subsessile or shortly petiolate;

blade elliptic to spatulate, oblanceolate, oblong, or obovate, 5–30(–35) × 2–8(–12) mm, base cuneate to attenuate, margin entire or irregularly dentate, densely scurfy abaxially, usually green and glabrate adaxially.

many, sessile or short petiolate;

blade obovate, oblong, oblanceolate, or elliptic, (5–)7–20 × 1–3(–5) mm, base cuneate, margin entire, apex acute.

Staminate flowers

with 3–5-parted perianth.

in glomerules in distal axils and short terminal spikes.

Pistillate flowers

intermixed with staminate in small axillary clusters.

in small axillary clusters.

Seeds

brown, 1–1.5 mm wide.

brown, 1.3–1.5 mm.

Fruiting

bracteoles subsessile or short stipitate, orbiculate, strongly compressed, 2–4 mm and as wide, united except at thin margin, margin dentate, terminal teeth often prominent, faces smooth or with cristate appendages (thornberi phase).

bracteoles sessile or subsessile, broadly obovate, 2–3 mm and as broad or about as broad, united 1/2 of length, margin free, deeply and sharply dentate, narrowed at summit, faces smooth or sometimes tuberculate.

Atriplex elegans

Atriplex coulteri

Phenology Flowering spring–fall.
Habitat Somewhat alkaline or clay low places, valley grasslands, coastal sage scrub, coastal slopes
Elevation 0-500 m (0-1600 ft)
Distribution
from FNA
AZ; CA; NM; NV; TX; UT; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Atriplex coulteri is closely allied to the geographically disjunct A. fruticulosa, from which it is said to differ in the compressed, small (2.5–3 mm) versus thickened and larger (3–5 mm) bracts. Specimens of A. fruticulosa, including the type, examined by me have bracteoles compressed-thickened, but hardly “globoid” as stated in the key to the species by H. M. Hall and F. E. Clements (1923). Additional specimens borrowed from California might clarify the situation; otherwise the two species are sufficiently close as to be treated as a single entity.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruiting bracteole margin dentate to incised, teeth 0.5-1 mm; bracteole not especially samaralike; s California e to Texas, and Mexico (Chihuahua, Sonora)
var. elegans
1. Fruiting bracteole margin finely toothed, teeth 0.3-0.5 mm; bracteole appearing samaralike; s Cali- fornia and w Arizona
var. fasciculata
Source FNA vol. 4, p. 364. FNA vol. 4, p. 363.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. elegans var. elegans, A. elegans var. fasciculata
Synonyms Obione elegans Obione coulteri
Name authority (Moquin-Tandon) D. Dietrich: Syn. Pl. 5: 537. (1852) (Moquin-Tandon) D. Dietrich: Syn. Pl. 5: 537. (1852)
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