Atriplex covillei |
Atriplex phyllostegia |
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Coville's orach |
leafcover saltweed, Truckee orach |
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Habit | Herbs, spreading, 1–4(–5) dm and as broad. | Herbs monoecious (or entirely pistillate), erect, much-branched, rounded, bushy, 0.5–6 dm. |
Stems | terete, sparsely scurfy when young. |
mostly ascending, terete. |
Leaves | petiole to 1/2 as long as blade (becoming subsessile distally); blade green or finally stramineous, (10–)20–50 × 6–30 mm, firm, base abruptly acute to narrowly cuneate, apex acute to attenuate, sparsely scurfy. |
petiole 0.3–2 cm; blade 10–50 × 5–25 mm, base varying from shortly hastate to truncate or cuneate. |
Staminate flowers | in sessile glomerules in distal axils, often mixed with pistillate ones, staminate calyx deeply 5-cleft; lobes obtuse, not appendaged. |
in small to moderate, axillary glomerules near ends of branches, or in bracteate terminal spikes. |
Pistillate flowers | with calyx of (1–)3(–5) hyaline sepals. |
in axillary clusters. |
Seeds | dark reddish brown, 1–1.5 mm. |
brown, 1.2 mm. |
Fruiting | bracteoles sessile or stipitate, 6–12 × 4–7 mm, margin mostly 3-lobed, with elongate terminal lobe triangular to lanceolate, 2 short rounded lobes at base or sides merely rounded at base, united to beyond middle. |
bracteoles lanceolate or oblong, 5–14(–20) mm, sharply lobed to tuberculate at base, also often sharply cristate below usually attenuate apical lobe; tips widely recurved-spreading. |
Atriplex covillei |
Atriplex phyllostegia |
|
Phenology | Flowering summer–fall. | Flowering Apr–Aug. |
Habitat | Mixed saltbush-greasewood, rabbitbrush, warm desert shrub, and salt grass communities in saline substrates | Valley bottoms, silty or clay, less commonly, sandy alluvium with greasewood |
Elevation | 800-1700 m (2600-5600 ft) | 1200-1500 m (3900-4900 ft) |
Distribution |
CA; NV; OR
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NV
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Discussion | Endolepis covillei was treated within the synonymy of Atriplex phyllostegia (Torrey) S. Watson by H. M. Hall and F. E. Clements (1923). H. C. Stutz et al. (1993) placed Atriplex covillei within Endolepis, based in large part on the presence of a perianth subtending the pistil within the fruiting bracteoles and on the lack of Kranz anatomy in the leaves. The pattern of venation is, nevertheless, very similar to that in species with Kranz anatomy. The presence of perianth scales in the pistillate flowers of A. covillei has been regarded as evidence of relationship with A. suckleyi. Despite placement of these taxa within Endolepis by Stutz et al. Atriplex covillei is possibly more closely allied to the morphologically similar and partially sympatric A. phyllostegia than it is to strongly dissimilar and the distantly disjunct A. suckleyi. Stutz and his associates placed great emphasis on the presence of reduced perianth segments subtending the pistil within the fruiting bracteoles of A. covillei. Calyces per se, otherwise known only in A. suckleyi and A. pleiantha, probably have arisen independently. Their presence does not necessarily indicate a close relationship. Stutz et al. pointed to other differences aside from the calyx of the pistillate flowers, and it is apparent that the two entities can stand as distinct species. To segregate A. covillei within a separate genus and to ally it with a species to which its relationships are obscure at best, stretches logic beyond reason. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 4. | FNA vol. 4, p. 368. |
Parent taxa | Chenopodiaceae > Atriplex > subg. Obione > sect. Covilleiae | Chenopodiaceae > Atriplex > subg. Obione > sect. Phyllostegiae |
Sibling taxa | ||
Synonyms | Endolepis covillei | Obione phyllostegia, A. draconis, A. phyllostegia var. draconis |
Name authority | (Standley) J. F. Macbride: Contr. Gray Herb. 53: 11. (1918) | (Torrey ex S. Watson) S. Watson: Proc. Amer. Acad. Arts 9: 108. (1874) |
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