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Coville's orach

Parry's saltbush

Habit Herbs, spreading, 1–4(–5) dm and as broad. Shrubs, dioecious, mainly 2–5 dm, armed.
Stems

terete, sparsely scurfy when young.

Leaves

petiole to 1/2 as long as blade (becoming subsessile distally);

blade green or finally stramineous, (10–)20–50 × 6–30 mm, firm, base abruptly acute to narrowly cuneate, apex acute to attenuate, sparsely scurfy.

, short petiolate to sessile;

blade orbiculate-cordate, or ovate-deltate to elliptic, 7–16(–22) × 6–10(–16) mm, base truncate or tapering, entire or less commonly subhastate.

Staminate flowers

in sessile glomerules in distal axils, often mixed with pistillate ones, staminate calyx deeply 5-cleft;

lobes obtuse, not appendaged.

in glomerules on interrupted, leafy paniculate spikes.

Pistillate flowers

with calyx of (1–)3(–5) hyaline sepals.

in small glomerules borne in leafy, paniculate spikes.

Seeds

dark reddish brown, 1–1.5 mm.

brown, 1.3–1.9 mm.

Fruiting

bracteoles sessile or stipitate, 6–12 × 4–7 mm, margin mostly 3-lobed, with elongate terminal lobe triangular to lanceolate, 2 short rounded lobes at base or sides merely rounded at base, united to beyond middle.

bracteoles sessile or nearly so, truncate-flabelliform, 3–4 × 3–5 mm, thick and rigid, spongy, united to beyond middle, margin entire, faces smooth.

Atriplex covillei

Atriplex parryi

Phenology Flowering summer–fall. Flowering spring–fall.
Habitat Mixed saltbush-greasewood, rabbitbrush, warm desert shrub, and salt grass communities in saline substrates Saline, fine-textured soils
Elevation 800-1700 m (2600-5600 ft) 60-1500 m (200-4900 ft)
Distribution
from FNA
CA; NV; OR
from FNA
CA; NV
[WildflowerSearch map]
[BONAP county map]
Discussion

Endolepis covillei was treated within the synonymy of Atriplex phyllostegia (Torrey) S. Watson by H. M. Hall and F. E. Clements (1923).

H. C. Stutz et al. (1993) placed Atriplex covillei within Endolepis, based in large part on the presence of a perianth subtending the pistil within the fruiting bracteoles and on the lack of Kranz anatomy in the leaves. The pattern of venation is, nevertheless, very similar to that in species with Kranz anatomy. The presence of perianth scales in the pistillate flowers of A. covillei has been regarded as evidence of relationship with A. suckleyi. Despite placement of these taxa within Endolepis by Stutz et al. Atriplex covillei is possibly more closely allied to the morphologically similar and partially sympatric A. phyllostegia than it is to strongly dissimilar and the distantly disjunct A. suckleyi. Stutz and his associates placed great emphasis on the presence of reduced perianth segments subtending the pistil within the fruiting bracteoles of A. covillei. Calyces per se, otherwise known only in A. suckleyi and A. pleiantha, probably have arisen independently. Their presence does not necessarily indicate a close relationship. Stutz et al. pointed to other differences aside from the calyx of the pistillate flowers, and it is apparent that the two entities can stand as distinct species. To segregate A. covillei within a separate genus and to ally it with a species to which its relationships are obscure at best, stretches logic beyond reason.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Atriplex parryi occurs with greasewood, saltbush, saltgrass, and Nitrophila. The cordate-clasping, sessile leaves and spiny branches distinguish this species from its nearest congeners.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 4. FNA vol. 4, p. 378.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Covilleiae Chenopodiaceae > Atriplex > subg. Pterochiton
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Synonyms Endolepis covillei
Name authority (Standley) J. F. Macbride: Contr. Gray Herb. 53: 11. (1918) S. Watson: Proc. Amer. Acad. Arts 17: 378. (1882)
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