Atriplex covillei
Atriplex holocarpa
Coville's orach
pop saltbush
terete, sparsely scurfy when young.
branching, diffuse or procumbent, softly scurfy-tomentose.
petiole to 1/2 as long as blade (becoming subsessile distally);
blade green or finally stramineous, (10–)20–50 × 6–30 mm, firm, base abruptly acute to narrowly cuneate, apex acute to attenuate, sparsely scurfy.
alternate;
petiole to 1/2 as long as blade;
blade obovate or rhombic to deltoid, 10–30 mm, base obtuse, margin sinuate to serrate, apex irregularly toothed, acute.
in axillary glomerules, staminate in distal axils surrounded by pistillate flowers, these only and usually few together in most axils, very small and globular at anthesis.
in sessile glomerules in distal axils, often mixed with pistillate ones, staminate calyx deeply 5-cleft;
lobes obtuse, not appendaged.
with calyx of (1–)3(–5) hyaline sepals.
dark reddish brown, 1–1.5 mm.
broadly elliptic;
radical lateral, erect.
bracteoles sessile or stipitate, 6–12 × 4–7 mm, margin mostly 3-lobed, with elongate terminal lobe triangular to lanceolate, 2 short rounded lobes at base or sides merely rounded at base, united to beyond middle.
bracteoles sessile, obovoid-globular, fused, scarcely compressed, 8–12 mm, of loosely fibrous and spongy consistency, with thin membranous epidermis and thin, inner membrane, opening at summit closed by 2 erect, appressed, entire or 3-toothed valves, apex shortly apiculate, not flattened at top.
Atriplex covillei
Atriplex holocarpa
Endolepis covillei was treated within the synonymy of Atriplex phyllostegia (Torrey) S. Watson by H. M. Hall and F. E. Clements (1923).
H. C. Stutz et al. (1993) placed Atriplex covillei within Endolepis, based in large part on the presence of a perianth subtending the pistil within the fruiting bracteoles and on the lack of Kranz anatomy in the leaves. The pattern of venation is, nevertheless, very similar to that in species with Kranz anatomy. The presence of perianth scales in the pistillate flowers of A. covillei has been regarded as evidence of relationship with A. suckleyi. Despite placement of these taxa within Endolepis by Stutz et al. Atriplex covillei is possibly more closely allied to the morphologically similar and partially sympatric A. phyllostegia than it is to strongly dissimilar and the distantly disjunct A. suckleyi. Stutz and his associates placed great emphasis on the presence of reduced perianth segments subtending the pistil within the fruiting bracteoles of A. covillei. Calyces per se, otherwise known only in A. suckleyi and A. pleiantha, probably have arisen independently. Their presence does not necessarily indicate a close relationship. Stutz et al. pointed to other differences aside from the calyx of the pistillate flowers, and it is apparent that the two entities can stand as distinct species. To segregate A. covillei within a separate genus and to ally it with a species to which its relationships are obscure at best, stretches logic beyond reason.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
I have seen no specimens of this species and therefore it is not mapped. H. M. Hall and F. E. Clements (1923) in discussion of the related Atriplex lindleyi (as A. halimoides) noted that it has “been grown in American gardens with the thought of using them as forage plants, but…has [not] been found suitable for general planting. P. G. Wilson (1984) indicated that the species is relatively widespread in Australia, mainly in southern parts, where it grows “often on flood-plains or sandy flats.”
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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