Atriplex coulteri
Atriplex suckleyi
Coulter's orach, Coulter's orache, Coulter's saltbush
Suckley's orach
frequently tinged with red, much branched, sparsely scurfy.
terete, often tinged with red, sparsely mealy or glabrous.
many, sessile or short petiolate;
blade obovate, oblong, oblanceolate, or elliptic, (5–)7–20 × 1–3(–5) mm, base cuneate, margin entire, apex acute.
numerous, alternate, sessile;
blade lanceolate to elliptic or less commonly ovate, 7–35 × (2–)4–10(–11) mm, thick and succulent, base acute, margin entire, apex acute or acuminate, glaucous, sparsely farinose when young.
in glomerules in distal axils and short terminal spikes.
in small glomerules, these in distal axils or in short, dense or interrupted, mostly simple, terminal spikes, perianth cup-shaped, frequently pinkish, lobes each with fleshy crest.
in small axillary clusters.
solitary or few in leaf axils, calyx within bracteoles of 3 or 4, entire or lobed, distinct, obtuse, hyaline sepals.
brown, 1.3–1.5 mm.
brown, 1.5 × 1.2 mm.
bracteoles sessile or subsessile, broadly obovate, 2–3 mm and as broad or about as broad, united 1/2 of length, margin free, deeply and sharply dentate, narrowed at summit, faces smooth or sometimes tuberculate.
bracteoles small and difficult to find, sessile, ovate, 2 × 1.5 mm, membranous, united to apex, without appendages, scurfy, each pistil subtended by a perianth.
Atriplex coulteri
Atriplex suckleyi
Of conservation concern.
Atriplex coulteri is closely allied to the geographically disjunct A. fruticulosa, from which it is said to differ in the compressed, small (2.5–3 mm) versus thickened and larger (3–5 mm) bracts. Specimens of A. fruticulosa, including the type, examined by me have bracteoles compressed-thickened, but hardly “globoid” as stated in the key to the species by H. M. Hall and F. E. Clements (1923). Additional specimens borrowed from California might clarify the situation; otherwise the two species are sufficiently close as to be treated as a single entity.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
At specific rank, the epithet suckleyi clearly has priority. Stutz et al. (1993) resurrected Endolepis Torrey and placed within it two completely disparate species, E. dioica and E. covillei Standley [Atriplex covillei (Standley) J. F. Macbride]. The interpretation of the genus by Stutz et al. stands on the presence of perianth segments subtending the ovary within the fruiting bracteoles, lack of Kranz leaf anatomy, and other more equivocal characteristics. The shared features hardly indicate near affinity, however. The two taxa are otherwise grossly dissimilar. Fundamentally, the genus Endolepis as resurrected by H. C. Stutz et al. stands on the basis of a single character: the presence of perianth segments. Kranz anatomy rises and falls, both within subg. Atriplex and subg. Obione. Thus, coincidence of the non-Kranz criterion is subject to interpretation. Perianth segments subtending the ovary within the enclosing bracteoles, per se, appear to be of independent origin. And, the peculiar nature of the staminate calyx in A. suckleyi (a major determining condition in establishment of the genus Endolepis by Torrey) is not present in A. covillei. Certainly the two species included within Endolepis appear to be as closely allied to other species of Atriplex as they are to each other.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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