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Coulter's orach, Coulter's orache, Coulter's saltbush

crownscale, wedgescale

Habit Herbs, perennial, sometimes flowering as an annual, spreading 0.7–10 dm, slightly woody at base. Herbs, erect or decumbent, 0.5–6 dm; branches terete, fructiferous almost to base, scurfy when young.
Stems

frequently tinged with red, much branched, sparsely scurfy.

Leaves

many, sessile or short petiolate;

blade obovate, oblong, oblanceolate, or elliptic, (5–)7–20 × 1–3(–5) mm, base cuneate, margin entire, apex acute.

alternate, subsessile or proximal short petiolate;

blade ascending, oblong, oblong-ovate, or lanceolate to elliptic, 5–40 × 2–10 mm, thin, base acute to obtuse, margin entire, apex acute to acuminate.

Flowers

of both sexes in small axillary glomerules, 5-merous.

Staminate flowers

in glomerules in distal axils and short terminal spikes.

Pistillate flowers

in small axillary clusters.

Seeds

brown, 1.3–1.5 mm.

light brown to amber, 1–1.5 mm.

Fruiting

bracteoles sessile or subsessile, broadly obovate, 2–3 mm and as broad or about as broad, united 1/2 of length, margin free, deeply and sharply dentate, narrowed at summit, faces smooth or sometimes tuberculate.

bracteoles almost sessile, broadly cuneate, 2–5 × (2–)2.5–5 mm, united to summit, truncate at summit, sides smooth or obscurely tuberculate.

2n

= 36.

Atriplex coulteri

Atriplex coronata

Phenology Flowering spring–fall.
Habitat Somewhat alkaline or clay low places, valley grasslands, coastal sage scrub, coastal slopes
Elevation 0-500 m (0-1600 ft)
Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Of conservation concern.

Atriplex coulteri is closely allied to the geographically disjunct A. fruticulosa, from which it is said to differ in the compressed, small (2.5–3 mm) versus thickened and larger (3–5 mm) bracts. Specimens of A. fruticulosa, including the type, examined by me have bracteoles compressed-thickened, but hardly “globoid” as stated in the key to the species by H. M. Hall and F. E. Clements (1923). Additional specimens borrowed from California might clarify the situation; otherwise the two species are sufficiently close as to be treated as a single entity.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 3 (3 in the flora).

Although placed within the Atriplex argentea species complex by P. C. Standley (1916), this taxon appears more nearly allied to A. cordulata, differing mainly in leaf shape, round-ovate or deltoid-ovate to elliptic or lanceolate (not ovate to cordate-ovate), and markedly tuberculate faces of the fruiting bracteoles.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems ± erect; body of fruiting bracteoles ± semicircular in profile, compressed or ± spheric; tubercles dense; marginal teeth ± equal.
var. notatior
1. Stems decumbent to ascending; body of fruiting bracteoles approaching circular or flabellate in profile, ± compressed; tubercles usually few; marginal teeth unequal
→ 2
2. Fruiting bracteoles 2-4 mm; basal branches and leaves typically opposite.
var. vallicola
2. Fruiting bracteoles (2-)4-5 mm; basal branches and leaves typically alternate
var. coronata
Source FNA vol. 4, p. 363. FNA vol. 4.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Pusillae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. coronata var. coronata, A. coronata var. notatior, A. coronata var. vallicola
Synonyms Obione coulteri A. elegans var. coronata, Obione coronata
Name authority (Moquin-Tandon) D. Dietrich: Syn. Pl. 5: 537. (1852) S. Watson: Proc. Amer. Acad. Arts 9: 114. (1874)
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