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crownscale, wedgescale

heart-leaf orach, heartscale

Habit Herbs, erect or decumbent, 0.5–6 dm; branches terete, fructiferous almost to base, scurfy when young. Herbs, erect, simple, oppositely or alternately branched from base and sparingly so distally, rigid, scurfy, 1–5 dm.
Stems

slender to rather coarse, initially erect-ascending, finally spreading-ascending, scurfy when young.

Leaves

alternate, subsessile or proximal short petiolate;

blade ascending, oblong, oblong-ovate, or lanceolate to elliptic, 5–40 × 2–10 mm, thin, base acute to obtuse, margin entire, apex acute to acuminate.

alternate except proximalmost ones, blade ovate to cordate-ovate, 5–15(–20) × 3–10(–13) mm;

base mostly cordate (to rounded), margin entire or denticulate, apex acute to attenuate, thickish and scurfy-tomentose.

Flowers

of both sexes in small axillary glomerules, 5-merous.

of both sexes mixed in small axillary clusters.

Staminate flowers

in glomerules in distal axils, 4–5-merous.

Seeds

light brown to amber, 1–1.5 mm.

deep red-brown, 1.5–1.8 mm.

Fruiting

bracteoles almost sessile, broadly cuneate, 2–5 × (2–)2.5–5 mm, united to summit, truncate at summit, sides smooth or obscurely tuberculate.

bracteoles sessile or subsessile, round-ovate (semi-orbiculate) to deltoid-rhombic or flabellate, slightly compressed, 3–5 mm and as broad, united to middle, deeply toothed, margin with acute teeth, terminal tooth subequal to others or largest, thin and soft at margin, hard at center, scarcely though sometimes tuberculate on faces.

2n

= 36.

= 36, 54.

Atriplex coronata

Atriplex cordulata

Distribution
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Although placed within the Atriplex argentea species complex by P. C. Standley (1916), this taxon appears more nearly allied to A. cordulata, differing mainly in leaf shape, round-ovate or deltoid-ovate to elliptic or lanceolate (not ovate to cordate-ovate), and markedly tuberculate faces of the fruiting bracteoles.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Stems ± erect; body of fruiting bracteoles ± semicircular in profile, compressed or ± spheric; tubercles dense; marginal teeth ± equal.
var. notatior
1. Stems decumbent to ascending; body of fruiting bracteoles approaching circular or flabellate in profile, ± compressed; tubercles usually few; marginal teeth unequal
→ 2
2. Fruiting bracteoles 2-4 mm; basal branches and leaves typically opposite.
var. vallicola
2. Fruiting bracteoles (2-)4-5 mm; basal branches and leaves typically alternate
var. coronata
1. Anthers red or purple; fruiting bracteoles 4-5 mm, central tooth largest; wc California
var. cordulata
1. Anthers yellow; fruiting bracteoles 3-3.5 mm, central tooth subequal to lateral; Kern, Kings, and Tulare counties, California
var. erecticaulis
Source FNA vol. 4. FNA vol. 4, p. 358.
Parent taxa Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Pusillae Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Pusillae
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. coronata var. coronata, A. coronata var. notatior, A. coronata var. vallicola
A. cordulata var. cordulata, A. cordulata var. erecticaulis
Synonyms A. elegans var. coronata, Obione coronata Obione cordulata
Name authority S. Watson: Proc. Amer. Acad. Arts 9: 114. (1874) Jepson: Pittonia 2: 304. (1892)
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