Atriplex coronata
Atriplex californica
crownscale, wedgescale
California orach, California saltbush
many branched, subterete, 1.5–5 dm, white scurfy when young.
alternate, subsessile or proximal short petiolate;
blade ascending, oblong, oblong-ovate, or lanceolate to elliptic, 5–40 × 2–10 mm, thin, base acute to obtuse, margin entire, apex acute to acuminate.
numerous, often crowded, alternate or proximalmost opposite;
blade narrowly lanceolate to narrowly oblanceolate or elliptic, 4–20 × 1–5 mm, acute at both ends, gray scurfy.
of both sexes in small axillary glomerules, 5-merous.
in terminal bracteate spikes, or mixed with pistillate in rather dense axillary clusters, 4-merous.
light brown to amber, 1–1.5 mm.
dark (black), 1–2 mm.
bracteoles almost sessile, broadly cuneate, 2–5 × (2–)2.5–5 mm, united to summit, truncate at summit, sides smooth or obscurely tuberculate.
bracteoles sessile, rhombic-ovate to ovate, scarcely united, 2.5–4 mm, margin entire, acute.
= 36.
Atriplex coronata
Atriplex californica
Varieties 3 (3 in the flora).
Although placed within the Atriplex argentea species complex by P. C. Standley (1916), this taxon appears more nearly allied to A. cordulata, differing mainly in leaf shape, round-ovate or deltoid-ovate to elliptic or lanceolate (not ovate to cordate-ovate), and markedly tuberculate faces of the fruiting bracteoles.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
H. M. Hall and F. E. Clements (1923) placed great emphasis on the inferior radicle, dioecious habit, and free bracts in stating that there are no close relatives of Atriplex californica in North America. P. C. Standley (1916) regarded the radicle as lateral or superior, not inferior, and placed the species at the beginning of his treatment of the American species. Plants of A. californica, however, form a mirror-image, matched pair with A. watsonii, with which they share habit, leaf conformation, staminate glomerules arranged, at least partially, in terminal interrupted spikes, and Kranz anatomy, but from which they differ in the radicle position, valves of the fruiting bracteoles free beyond the middle, monoecious habit, and mostly alternate leaves. The interpretation by Hall and Clements of radicle position as fundamental is made problematic by the apparent random occurrence of a great many morphologic features from place to place within the genus and often within the taxon. Each character fails individually as having definitive taxonomic importance, making difficult or impossible an ultimately satisfactory phylogenetic arrangement. It is not, however, the character that makes the species, rather, it is the entire syndrome of features that constitutes the taxon. Most certainly A. californica is more closely allied to American taxa than to those from other regions of the world.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Stems ± erect; body of fruiting bracteoles ± semicircular in profile, compressed or ± spheric; tubercles dense; marginal teeth ± equal. | var. notatior |
1. Stems decumbent to ascending; body of fruiting bracteoles approaching circular or flabellate in profile, ± compressed; tubercles usually few; marginal teeth unequal | → 2 |
2. Fruiting bracteoles 2-4 mm; basal branches and leaves typically opposite. | var. vallicola |
2. Fruiting bracteoles (2-)4-5 mm; basal branches and leaves typically alternate | var. coronata |
CA
WildflowerSearch
iNaturalist
USDA Plants Database
