Atriplex argentea |
Atriplex mucronata |
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maidenhair spleenwort, silver orach, silver orache, silver saltbush, silverscale, silverscale orache, silverscale saltbush, silvery orache |
quelite |
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Habit | Herbs, simple or freely branched, 0.5–6 dm; branches rather stout, angled, scurfy when young. | Herbs, annual. | ||||||||||||||||
Stems | erect, ascending, or procumbent, much branched, obtusely angled, 1–6 dm, stout, scurfy when young. |
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Leaves | often opposite proximally, petiolate or distal bracteate ones subsessile, blade lance-ovate, lanceolate, deltoid, or cordate, 5–75 × 4–50(–75) mm, base subhastate or obtuse to acute, margin entire or essentially so, sometimes closely repand-dentate, apex obtuse to acute or rounded, scurfy (glabrous). |
alternate or proximalmost opposite or subopposite; petiole to 1.5 cm or sessile; blade paler abaxially, oblong or oval, lanceolate, or elliptic to broadly obovate or narrowly oblong, 10–40(–60) × (2–)4–20 mm, base rounded to cuneate, margin entire or undulate, rarely with 1–2 teeth, apex rounded to acute, mucronate, thin, often densely white scurfy beneath, grayish green and glabrate above. |
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Flowers | in axillary glomerules and terminal, interrupted spikes. |
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Staminate flowers | borne in distal axils, or in short dense spikes or panicles, or intermixed with pistillate, with 4–5-parted calyx. |
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Pistillate flowers | fascicled in axils. |
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Seeds | brown, 1.5–2 mm wide; radicle superior or lateral. |
reddish brown, 2 mm. |
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Fruiting | bracteoles sessile, subsessile, or stipitate (stipe 0.5–5 mm), cuneate-orbicular, (2.5–)4–11.2 × 2–8.8(–14) mm, margin foliaceous below apex, subentire or dentate to laciniate, face smooth, tuberculate, or crested, processes sometimes again toothed, teeth then aligned with axis of process. |
bracteoles subsessile or with stipes to 1 mm, cuneate-orbiculate, compressed, 4.5–7 × 3.5–5.6 mm, typically longer than wide, united to middle, apex rounded, 3–5-toothed, teeth subequal, sides irregularly tuberculate or with 2 lateral dentate crests, rarely not appendaged. |
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Staminate | glomerules terminal or in dense or interrupted, terminal or axillary, naked spikes or shortly branched panicles. |
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2n | = 18, 36, 54. |
= 18. |
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Atriplex argentea |
Atriplex mucronata |
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Phenology | Flowering summer–fall. | |||||||||||||||||
Habitat | Sandy seashores, salt marshes | |||||||||||||||||
Elevation | 0 m (0 ft) | |||||||||||||||||
Distribution |
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB; SK; Mexico
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AL; FL; LA; MA; MD; MS; NC; NH; NJ; TX; VA |
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Discussion | Varieties 5 (5 in the flora). Herbarium materials have tended to represent a catchall for annual specimens not readily assignable to other taxa. Indeed, the distinguishing features of the Atriplex argentea complex are shared singly and often in combination with other taxa. Only by use of combinations of features can this taxon be defined. Those features, with much variation, center around the broad, typically ovate to deltoid leaf blades (often definitely 3-veined) and more-or-less compressed, sessile to subsessile (or short stipitate), fruiting bracteoles on which the marginal processes, or teeth, are mainly aligned with the plane compression, and with the faces quite smooth to variously appendaged. Still some specimens are apparently intermediate with other species, especially with the closely allied A. saccaria, with which it is at least partially sympatric. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Problems with prior applications of the name Atriplex mucronata lie in the mistaken determination of the place of publication as Amer. Monthly Mag. & Crit. Rev. 2: 176. 1818 (where the name is only mentioned), instead of 2: 119. 1817 (where accompanied by a description and notes). H. M. Hall and F. E. Clements (1923) evidently relied on P. C. Standley’s (1916) interpretation, which indicated the wrong citation (see also A. dioica Rafinesque as an identical example). Hall and Clements applied the name mucronata to their interpretation of A. patula subsp. hastata or to A. hastata (i.e., to A. prostrata according to this treatment). The treatment of Obione by C. H. B. A. Moquin-Tandon (1849) included “A. mucronata Rafin.!” as a synonym of O. arenaria. Possibly a sheet in the Prodromus herbarium at Geneva was the basis for that decision. It has two mounted specimens, one labeled A. arenaria Nuttall, collected by Nuttall in “N. Jersey, 1826,” and a second labeled “Atriplex mucronata Rafinesque (A. arenaria Nuttall, N. Jersey) Maritime NY, Rafinesque 1819.” From that information (although the year is 1819, not 1817), and from the description of the taxon, it seems clear that A. arenaria Nuttall is a later synonym of A. mucronata Rafinesque by at least half a year. H. A. Gleason and A. Cronquist (1991) indicated that this taxon, by whatever name, is “perhaps better treated as a variety of the more tropical Atriplex pentandra (Jacquin) Standley, but the proper nomenclatural combination not yet made.” The present writer agrees with that conclusion, but such subjugation might indicate further contraction into the species of additional closely related taxa, e.g., A. wrightii, which is clearly closely allied as well. Plants from the coastal states from New England south to New Jersey are much alike and seldom, if ever, display prominent, terminal, naked spikes or panicles with beadlike glomerules of staminate flowers. Plants from Florida westward sometimes have such staminate spikes or panicles. Specimens from Florida and some from Texas can be distinguished from Atriplex pentandra only with difficulty, especially those individuals with entire leaves. However, the fruiting bracteole length-width ratio and overall shape, with some allowance for overlap, can serve to distinguish most specimens; those of A. mucronata are proportionately longer than broad and, on average, larger. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4. | FNA vol. 4, p. 362. | ||||||||||||||||
Parent taxa | Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Argenteae | Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Arenariae | ||||||||||||||||
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Synonyms | Obione argentea | A. arenaria, A. cristata var. arenaria, A. pentandra subsp. arenaria | ||||||||||||||||
Name authority | Nuttall: Gen. N. Amer. Pl. 1: 198. (1818) | Rafinesque: Amer. Monthly Mag. & Crit. Rev. 2(2): 119. (1817) | ||||||||||||||||
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