Atriplex argentea |
Atriplex joaquiniana |
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maidenhair spleenwort, silver orach, silver orache, silver saltbush, silverscale, silverscale orache, silverscale saltbush, silvery orache |
San Joaquin orach |
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Habit | Herbs, simple or freely branched, 0.5–6 dm; branches rather stout, angled, scurfy when young. | Herbs, monoecious or subdioecious, erect, sparsely branched, (1–)3–10 dm; branches obtusely angled, rather rigidly ascending, finely farinose when young. | ||||||||||||||||
Leaves | often opposite proximally, petiolate or distal bracteate ones subsessile, blade lance-ovate, lanceolate, deltoid, or cordate, 5–75 × 4–50(–75) mm, base subhastate or obtuse to acute, margin entire or essentially so, sometimes closely repand-dentate, apex obtuse to acute or rounded, scurfy (glabrous). |
mostly alternate; petiole 0.5–2.5 cm, becoming shorter or subsessile in distal ones; blade deltoid to rhombic-ovate or lanceolate, (10–)15–50(–70) × 8–40 mm, base rounded, truncate, or broadly cuneate, margin irregularly sinuate-dentate or repand-dentate to entire, sometimes subhastate, apex obtuse to acute; distal blades often narrower and sometimes entire except for basal lobes. |
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Flowers | in axillary glomerules and terminal, interrupted spikes. |
in dense or interrupted, naked, simple or paniculate spikes mainly 5–8 mm thick, staminate ones 4-merous. |
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Staminate flowers | borne in distal axils, or in short dense spikes or panicles, or intermixed with pistillate, with 4–5-parted calyx. |
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Seeds | brown, 1.5–2 mm wide; radicle superior or lateral. |
dark brown or black, 0.8–1.5 mm; radicle inferior. |
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Fruiting | bracteoles sessile, subsessile, or stipitate (stipe 0.5–5 mm), cuneate-orbicular, (2.5–)4–11.2 × 2–8.8(–14) mm, margin foliaceous below apex, subentire or dentate to laciniate, face smooth, tuberculate, or crested, processes sometimes again toothed, teeth then aligned with axis of process. |
bracteoles densely packed on rachis, sessile, ovate-oblong or rounded-deltoid, (2–)2.5–3(–4.5) mm, united only at rounded or truncate base, angled and cristate on faces, thin or spongy-thickened. |
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2n | = 18, 36, 54. |
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Atriplex argentea |
Atriplex joaquiniana |
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Phenology | Flowering summer. | |||||||||||||||||
Habitat | Alkali sink scrub or alkaline grasslands | |||||||||||||||||
Elevation | 0-200(-300) m (0-700(-1000) ft) | |||||||||||||||||
Distribution |
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; BC; MB; SK; Mexico
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CA |
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Discussion | Varieties 5 (5 in the flora). Herbarium materials have tended to represent a catchall for annual specimens not readily assignable to other taxa. Indeed, the distinguishing features of the Atriplex argentea complex are shared singly and often in combination with other taxa. Only by use of combinations of features can this taxon be defined. Those features, with much variation, center around the broad, typically ovate to deltoid leaf blades (often definitely 3-veined) and more-or-less compressed, sessile to subsessile (or short stipitate), fruiting bracteoles on which the marginal processes, or teeth, are mainly aligned with the plane compression, and with the faces quite smooth to variously appendaged. Still some specimens are apparently intermediate with other species, especially with the closely allied A. saccaria, with which it is at least partially sympatric. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Atriplex joaquiniana commonly occurs with Distichlis spicata, Allenrolfea occidentalis, Suaeda moquinii, Frankenia salina, Hordeum depressum, Spergularia macrotheca, and various annual species. H. M. Hall and F. E. Clements (1923) placed the relationship of this very distinctive California endemic with the Atriplex patula complex, with which one can make out a distant affinity. It differs from all other members previously treated within that complex in the small, more or less quadrangular fruiting bracteoles, which are typically crested on one or both elevated faces and have entire or less commonly dentate margins. The bracteoles are borne in very compact simple or paniculate spikes. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 4. | FNA vol. 4, p. 335. | ||||||||||||||||
Parent taxa | Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Argenteae | Chenopodiaceae > Atriplex > subg. Atriplex > sect. Teutliopsis | ||||||||||||||||
Sibling taxa | ||||||||||||||||||
Subordinate taxa | ||||||||||||||||||
Synonyms | Obione argentea | A. spicata, A. patula subsp. spicata, A. spicata var. lagunita | ||||||||||||||||
Name authority | Nuttall: Gen. N. Amer. Pl. 1: 198. (1818) | A. Nelson: Proc. Biol. Soc. Wash. 17: 99. (1904) | ||||||||||||||||
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