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armed saltbush, burscale, tubercled saltbush

Bakersfield smallscale, Tulare orach, Tulare saltbush

Habit Shrubs or subshrubs, dioecious, evergreen, mainly 2–10 × 4–10+ dm, woody especially basally, unarmed; branchlets obtusely angled to subterete. Herbs, erect, simple or sparingly branched, 1.5–4(–10) dm, white scurfy.
Stems

aging red, terete or obscurely angled, brittle.

Leaves

persistent, proximal ones opposite, becoming alternate distally, short petiolate or subsessile;

blade oblong to oblong-lanceolate, ovate, obovate, or spatulate, 12–40(–50) × 5–25 mm, base commonly subhastate to cuneate, margin entire or sinuate-dentate to strongly undulate-crisped, apex acute.

alternate or proximalmost opposite;

blade lanceolate to ovate, 6–20 × 4–8 mm, base rounded, margin entire, apex acute to acuminate, gray scurfy.

Staminate flowers

in interrupted or crowded glomerules 2–4.5 mm thick, in sparsely leafy paniculate spikes to 5+ dm.

in small axillary dense glomerules, 4-merous.

Pistillate flowers

few to solitary, in axillary clusters or in crowded or interrupted, often leafy, erect, branched spicate racemes or racemose panicles to 25+ cm.

solitary or in small axillary clusters, below or mixed with staminate.

Seeds

brown, 1.5–2 mm.

dark brown, 1–1.2 mm.

Fruiting

bracteoles on slender or stout stipes (2–)4–20 mm (or sessile), body broadly elliptic to globose, 6–15 mm and wide, spongy, united to the linear apex, margin deeply laciniate, faces appendaged with flattened to hornlike tubercles to 8 mm.

bracteoles sessile, rhombic-ovate, acute or acuminate, 2.5–3.5 × 2.2–3 mm, united to middle, thin margin toothed, faces plane, scurfy.

Atriplex acanthocarpa

Atriplex tularensis

Phenology Flowering summer–fall.
Habitat Alkaline plains, edges of alkali sink
Elevation 90-200 m (300-700 ft)
Distribution
from FNA
AZ; NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
CA
[BONAP county map]
Discussion

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

H. M. Hall and F. E. Clements (1923) discussed the relationship of this plant to Atriplex cordulata, noting that, “A. tularensis is a much more slender plant, the leaves are narrower in proportion to their length,” and tapering to the base, never at all cordate. They noted further, that “The fruiting bracts are sometimes very similar in shape, but those of tularensis typically end in an abrupt acute tooth not present in cordulata and both bract and seed are always smaller.” Because of its small fruiting bracteoles it was placed within the Pusillae by P. C. Standley (1916). The erect stature, coupled with distinctive spreading-ascending and stiff (at least in dried material) leaves, and foliose bracts apparently distinguish A. tularensis from A. coronata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blade ovate to rhombic-ovate, elliptic, or spatulate, margin mainly sinuate-dentate; mature fruiting inflorescences ascending to erect; west Texas, s New Mexico, se Arizona
var. acanthocarpa
1. Leaf blade, at least some, hastate-lanceolate, or strongly undulate-crisped; mature fruiting inflorescences mainly horizontally spreading; s Texas, Mexico
var. coahuilensis
Source FNA vol. 4, p. 370. FNA vol. 4, p. 359.
Parent taxa Chenopodiaceae > Atriplex > subg. Pterochiton Chenopodiaceae > Atriplex > subg. Obione > sect. Obione > subsect. Pusillae
Sibling taxa
A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. acanthocarpa var. acanthocarpa, A. acanthocarpa var. coahuilensis
Synonyms Obione acanthocarpa A. cordulata var. tularensis, Obione tularensis
Name authority (Torrey) S. Watson: Proc. Amer. Acad. Arts 9: 117. (1874) Coville: Contr. U.S. Natl. Herb. 4: 182, plate 19. (1893)
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