The green links below add additional plants to the comparison table. Blue links lead to other Web sites.
enable glossary links

common smoothcap, common smoothcap moss, undulate atrichum moss

cylindrical atrichum moss

Habit Plants small to fairly large, dark green. Plants dark green.
Stems

2–5 cm.

(2.5–)4–5 cm, densely leafy.

Leaves

sparse below, more crowded above, 4–9 × 0.8–1.5 mm, lingulate to lanceolate, undulate, keeled distally, concave to flat proximally, with oblique rows of abaxial teeth on undulations, the apex acute;

costa percurrent, with teeth on abaxial surface in distal half;

lamellae 2–6, 2–4(–7) cells high;

median leaf cells 20–32 µm wide, irregularly angled to hexagonal, sometimes rounded, often transversely elongate, convex on both surfaces, smooth or with minute, verrucose or striate papillae on abaxial surface, walls firm, strongly collenchymatous with trigones.

7–8 × 0.8–1 mm wide midway between base and tip, narrowly triangular-lanceolate to linear, subfalcate, plane, not undulate, with a few scattered abaxial teeth, or none, long-tapering to a narrowly acute apex, margin stoutly bordered, doubly toothed from proximal 1/8 of leaf, often colored;

costa spinose abaxially towards leaf apex;

lamellae (4–)5(–6), low and strictly parallel, (1–)3–4 cells high;

median leaf cells 20–25 µm wide, rounded hexagonal, with their longest dimension at right angles to the costa, with distinct trigones, smooth or occasionally with striate papillae near base of leaf.

Seta

1(–3) per perichaetium, 1–3 cm, reddish brown.

reddish brown, 3–4(–5) cm, usually 1 per perichaetium, occasionally 2.

Sexual condition

polygamous, some shoots bisexual, others apparently male or female;

perigonial bracts broad, forming antheridial buds, often several buds per plant.

polygamous, most shoots apparently unisexual and female;

unisexual male shoots not reported, perigonial leaves similar to the others, but with broader bases, to broadly lanceolate and unbordered in proximal 1/2, the innermost bracts tiny, obovate.

Capsule

2–4(–8) × 0.5–1 mm, curved to distinctly arcuate, almost horizontal;

operculum 2–3 mm.

6–8(–10) × 0.8–1 mm, reddish brown, finely striate, narrowly cylindric, somewhat curved, erect to inclined.

Spores

(12–)16–28 µm.

18–20 µm.

Atrichum undulatum

Atrichum cylindricum

Phenology Capsules mature spring–summer (Apr- Jul).
Habitat Soil, dry weedy habitats, especially roadside ditches Moist soil, ravines, ditches, stream banks in mixed forest, low wet woods and bottomland (swamp forest)
Elevation usually low elevations low elevations
Distribution
from FNA
ME; BC; NB; NF; NS; ON; PE; Europe; w Asia; c Asia; n Africa [Introduced in North America]
[WildflowerSearch map]
from FNA
AL; FL; IL; KY; LA; MO; MS; NC; SC; TN; TX; VA
Discussion

No Atrichum species occurring in North America has been as widely misunderstood as A. undulatum. Evidently introduced from Europe, its weedy habitat and strongly arcuate, almost horizontal capsules are the surest means of recognition. Variety minus, a fixture of European manuals, is a stunted form that is highly variable and possibly a hybrid. Its occurrence in North America has not been demonstrated. As used by American authors (e.g., O. E. Jennings 1951), it probably refers to A. altecristatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Atrichum cylindricum is distributed along the Atlantic and Gulf coastal plain from Virginia to Florida and eastern Texas, and in the Mississippi Embayment and Cumberland Plateau. In addition to the published distribution (Gary L. Smith 1977), the species has also been found on hummocks in a cypress swamp in southern Illinois. Atrichum crispulum is similar in size to A. cylindricum, but is dioicous, with broader, distinctly undulate leaves and well-developed teeth on the back of the lamina. Atrichum altecristatum is a more slender plant with taller lamellae, shorter, broader, undulate leaves, toothed on the back and less strongly bordered, and shorter capsules.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 27, p. 154. FNA vol. 27, p. 153.
Parent taxa Polytrichaceae > Atrichum Polytrichaceae > Atrichum
Sibling taxa
A. altecristatum, A. angustatum, A. crispulum, A. crispum, A. cylindricum, A. flavisetum, A. selwynii, A. tenellum
A. altecristatum, A. angustatum, A. crispulum, A. crispum, A. flavisetum, A. selwynii, A. tenellum, A. undulatum
Synonyms Polytrichum undulatum Polytrichum cylindricum, A. undulatum var. attenuatum
Name authority (Hedwig) P. Beauvois: Prodr. Aethéogam., 42. (1805) (F. Weber) G. L. Smith: Brittonia 29: 371. (1977)
Web links