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Bigelow's woolly locoweed, woolly locoweed

purple locoweed, woolly locoweed, woolly milkvetch

Habit Plants shortly caulescent, robust. Plants densely or loosely tuft-forming, acaulescent to subacaulescent or shortly caulescent, (6–)10–30(–45) cm, villous-tomentose.
Stems

(0 or)3–17 cm.

usually few or several, sometimes single, decumbent to ascending, when developed, with several, short internodes, to 2 cm, villous-tomentose.

Leaves

9–26 cm;

stipules 6–20 mm;

leaflets (13–)19–27, blades ovate, obovate, oval, or broadly elliptic, 6–25 mm.

(2–)5–32 cm;

stipules (3–)4–20 mm;

leaflets (9 or)11–35, blades elliptic, suborbiculate, ovate, obovate, rhombic-elliptic, rhombic-obovate, rhombic-ovate, oblong-elliptic, oblanceolate, or oval, (2–)3–30(–45) mm, apex acute, obtuse, retuse, or subacute, surfaces villous-tomentose, sometimes densely so.

Racemes

somewhat densely (15–)20–45-flowered, flowers subcontiguous or interrupted proximally;

axis (4–)5–11 cm in fruit.

(5–)7–45-flowered;

axis (0.5–)1–18 cm in fruit;

bracts 2.5–10(–12) mm;

bracteoles 0(–2).

Peduncles

(5–)8–22 cm.

ascending, (1.5–)2.5–24 cm.

Pedicels

0.5–3 mm.

Flowers

calyx 10.5–13.5 cm, tube (8–)8.3–10.3 × (3.2–)4–5.2 mm, lobes (1.7–)2.6–4.4 mm;

corolla pink-purple;

banner 17–22.5 mm;

keel 13.5–18.5 mm.

11.8–25 mm;

calyx cylindric to deeply campanulate, 6.8–15.5 mm, villous-tomentose, tube (4.5–)5–13 mm, lobes lanceolate to subulate or subulate-setaceous, 1.6–5.5(–6.8) mm;

corolla pink-purple, pale purple, pinkish, cream, or yellowish suffused with dull lavender, or tipped and suffused or margined with dull purple;

keel 9–20.5 mm.

Legumes

gently incurved or ± straight, ovoid-acuminate or lanceoloid-ellipsoid, sometimes slightly turgid, 10–15 × (4–)4.5–8 mm, stiffly papery or leathery, densely villous-tomentulose, hairs to 1–1.6 mm;

beak bilocular.

spreading or ascending, ± straight to incurved through 0.25+ spiral, oblong-ellipsoid, lanceoloid-ellipsoid, ovoid, or ovoid-ellipsoid, sometimes decidedly inflated (not bladdery), terete when narrow, obcompressed when broad, (6–)9–24 × (3–)4–13 mm, fleshy, stiffly papery, leathery, or subligneous, glabrous, puberulent, hispidulous, villosulous, villous-hirsute, villous-tomentulose, or villous-tomentose;

beaks sometimes unilocular.

Seeds

20–31.

12–38[–41].

2n

= 22.

= 22.

Astragalus mollissimus var. bigelovii

Astragalus mollissimus

Phenology Flowering (Jan–)Mar–Jun.
Habitat Dry plains and foothills, in desert- or mesquite-grasslands, among junipers, on calcareous soils, sandy loams, basalt gravel, over-grazed and badly eroded cattle ranges.
Elevation 1200–1900(–2300) m. (3900–6200(–7500) ft.)
Distribution
from FNA
AZ; NM; TX
[BONAP county map]
from USDA
w United States; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Variety bigelovii is known from extreme western Texas to southeastern Arizona and in New Mexico as far north as Socorro County and the Plains of San Augustin in Catron County.

D. Isely (1998) recognized var. bigelovii at the species level, and included with it vars. marcidus, matthewsii, and mogollonicus, primarily on the basis of the completely bilocular fruits.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 10 (8 in the flora).

The Astragalus mollissimus complex has been variously interpreted as belonging to several species, or has been combined in varietal status under one or more species, or has been split into species each with segregate varieties, and with peripheral taxa still represented at specific rank (D. Isely 1998). Isely (1983) recognized three species within the complex, restricting A. mollissimus to those phases with glabrous or sparsely puberulent fruits, A. bigelovii to the bulk of the remainder with shaggy-hairy fruits, and with A. thompsoniae (whose fruit has a unilocular beak) standing by itself, remote from the remainder of A. bigelovii in the broad sense. Two varieties are exclusive to Mexico: var. irolanus (M. E. Jones) Barneby occurs from Durango and Nuevo León southward to Puebla, and var. nitens Barneby is known from Coahuila. The present treatment follows R. C. Barneby (1964). Some of the varieties, possibly all, are poisonous to domestic livestock, especially to horses (N. Ritter 1917; L. F. James and S. L. Welsh 1992; M. H. Ralphs et al. 2002).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Legumes glabrous or sparsely pubescent (hispid­ulous, villous-tomentose, or loosely strigulose).
→ 2
2. Corollas cream, immaculate; Edwards Plateau, Texas.
var. coryi
2. Corollas pinkish, pink-purple, pure pale yel­low or yellowish suffused with dull lavender, or tipped and suffused or margined with dull purple; not of Edwards Plateau, Texas.
→ 3
3. Calyx tubes 3.4–4.5 mm wide; plains from Nebraska to New Mexico and w Texas.
var. mollissimus
3. Calyx tubes 0.8–3(–3.2) mm wide; se New Mexico, trans-Pecos Texas, southward.
var. earlei
1. Legumes densely pubescent (villous-tomentose, villous-hirsute, or villous-tomentulose).
→ 4
4. Racemes (5–)7–12-flowered; peduncles (1.5–)2.5–8 cm; legumes broadly ovoid, 7–13 mm wide; e Arizona (Apache County), nw New Mexico (Santa Fe to McKinley County).
var. matthewsii
4. Racemes 7–45-flowered; peduncles 2.5–24 cm; legumes ovoid, lunately ellipsoid, or lanceoloid-ellipsoid, 3–8(–11) mm wide; Arizona, Colorado, Idaho, Nevada, New Mexico, Texas, Utah.
→ 5
5. Legumes ovoid, turgid, 6–11 mm wide, beaks unilocular; Arizona and New Mexico northward to Idaho.
var. thompsoniae
5. Legumes ovoid, lunately ellipsoid or lanceoloid-ellipsoid, usually solid, rarely slightly turgid, 3–8 mm wide, beaks biloc­ular; Arizona, New Mexico, Texas.
→ 6
6. Plants acaulescent; calyces 6.8–9.9 mm; banners 11.8–16.3 mm, keel 10.3–12.5 mm; legumes 6–10 mm; seeds 12–16; trans-Pecos Texas (Presidio and Jeff Davis counties).
var. marcidus
6. Plants acaulescent or shortly caulescent; calyces 10–15.3 mm; banners 16–22.5 mm, keel 12.3–18.6 mm; legumes 9–15 mm; seeds 19–31; Arizona, New Mexico, Texas.
→ 7
7. Plants usually robust, shortly caulescent, stems (0 or)3–17 cm; leaves 9–26 cm, leaflet blades 6–25 mm; keel 13.5–18.5 mm; legume hairs to 1–1.6 mm; se Arizona to extreme w Texas; 1200–1900(–2300) m.
var. bigelovii
7. Plants usually dwarf, acaulescent; stems reduced to thick crowns; leaves (4–)6–16 cm, leaflet blades 3–13 mm; keel 12.5–14.5 mm; legume hairs to 1.6–2.6 mm; plateaus of n Arizona and wc New Mexico, on and near Mogollon Escarpment or north; 1800–2300 m.
var. mogollonicus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Astragalus > sect. Mollissimi > Astragalus mollissimus Fabaceae > subfam. Faboideae > Astragalus > sect. Mollissimi
Sibling taxa
A. mollissimus var. coryi, A. mollissimus var. earlei, A. mollissimus var. marcidus, A. mollissimus var. matthewsii, A. mollissimus var. mogollonicus, A. mollissimus var. mollissimus, A. mollissimus var. thompsoniae
A. accidens, A. accumbens, A. ackermanii, A. acutirostris, A. adanus, A. aequalis, A. agnicidus, A. agrestis, A. albens, A. albulus, A. allochrous, A. alpinus, A. altus, A. alvordensis, A. amblytropis, A. americanus, A. amnis-amissi, A. amphioxys, A. ampullarioides, A. ampullarius, A. andersonii, A. anisus, A. anserinus, A. applegatei, A. aquilonius, A. aretioides, A. argophyllus, A. aridus, A. arizonicus, A. arrectus, A. arthurii, A. asclepiadoides, A. asotinensis, A. asymmetricus, A. atratus, A. atropubescens, A. atwoodii, A. austiniae, A. australis, A. barnebyi, A. barrii, A. beathii, A. beatleyae, A. beckwithii, A. bernardinus, A. bibullatus, A. bicristatus, A. bisulcatus, A. bodinii, A. bolanderi, A. bourgovii, A. brandegeei, A. brauntonii, A. brazoensis, A. breweri, A. californicus, A. callithrix, A. calycosus, A. camptopus, A. canadensis, A. caricinus, A. casei, A. castaneiformis, A. castetteri, A. ceramicus, A. cerussatus, A. chamaeleuce, A. chamaemeniscus, A. chinensis, A. chloödes, A. chuskanus, A. cibarius, A. cicer, A. cimae, A. claranus, A. clevelandii, A. cliffordii, A. cobrensis, A. coccineus, A. collinus, A. coltonii, A. columbianus, A. concordius, A. congdonii, A. conjunctus, A. consobrinus, A. contortuplicatus, A. convallarius, A. cottamii, A. crassicarpus, A. cremnophylax, A. cronquistii, A. crotalariae, A. curtipes, A. curvicarpus, A. cusickii, A. cutleri, A. cyaneus, A. cymboides, A. deanei, A. debequaeus, A. desereticus, A. desperatus, A. deterior, A. detritalis, A. diaphanus, A. didymocarpus, A. distortus, A. diversifolius, A. douglasii, A. drabelliformis, A. drummondii, A. duchesnensis, A. eastwoodiae, A. egglestonii, A. emoryanus, A. endopterus, A. ensiformis, A. episcopus, A. equisolensis, A. eremiticus, A. ertterae, A. eucosmus, A. eurekensis, A. eurylobus, A. falcatus, A. feensis, A. filipes, A. flavus, A. flexuosus, A. fucatus, A. funereus, A. gambelianus, A. geyeri, A. gibbsii, A. giganteus, A. gilensis, A. gilmanii, A. gilviflorus, A. glycyphyllos, A. gracilis, A. grayi, A. gypsodes, A. hallii, A. hamiltonii, A. harrisonii, A. heilii, A. henrimontanensis, A. holmgreniorum, A. hoodianus, A. hornii, A. howellii, A. humillimus, A. humistratus, A. hyalinus, A. hypoxylus, A. inflexus, A. insularis, A. inversus, A. inyoensis, A. iodanthus, A. iodopetalus, A. iselyi, A. jaegerianus, A. jejunus, A. johannis-howellii, A. kelseyae, A. kentrophyta, A. kerrii, A. knightii, A. laccoliticus, A. lancearius, A. laxmannii, A. layneae, A. leibergii, A. lemmonii, A. lentiformis, A. lentiginosus, A. leptaleus, A. leptocarpus, A. leucolobus, A. limnocharis, A. lindheimeri, A. linifolius, A. loanus, A. lonchocarpus, A. lotiflorus, A. lutosus, A. lyallii, A. macrodon, A. magdalenae, A. malacoides, A. malacus, A. megacarpus, A. michauxii, A. microcymbus, A. microcystis, A. microlobium, A. micromerius, A. miguelensis, A. minthorniae, A. misellus, A. miser, A. missouriensis, A. moencoppensis, A. mohavensis, A. molybdenus, A. monoensis, A. montii, A. monumentalis, A. mulfordiae, A. multiflorus, A. musiniensis, A. naturitensis, A. neglectus, A. nelsonianus, A. neomexicanus, A. nevinii, A. newberryi, A. nidularius, A. nothoxys, A. nudisiliquus, A. nutans, A. nutriosensis, A. nuttallianus, A. nuttallii, A. nutzotinensis, A. nyensis, A. obcordatus, A. obscurus, A. oniciformis, A. oocalycis, A. oocarpus, A. oophorus, A. oreganus, A. osterhoutii, A. oxyphysus, A. pachypus, A. palmeri, A. panamintensis, A. pardalinus, A. parryi, A. pattersonii, A. pauperculus, A. paysonii, A. peckii, A. pectinatus, A. perianus, A. phoenix, A. pictiformis, A. pinonis, A. piscator, A. piutensis, A. plattensis, A. platytropis, A. polaris, A. pomonensis, A. porrectus, A. praelongus, A. preussii, A. proimanthus, A. proximus, A. pseudiodanthus, A. pterocarpus, A. pubentissimus, A. pulsiferae, A. puniceus, A. purshii, A. pycnostachyus, A. racemosus, A. rafaelensis, A. rattanii, A. ravenii, A. recurvus, A. reflexus, A. remotus, A. reventiformis, A. reventus, A. riparius, A. ripleyi, A. robbinsii, A. rusbyi, A. sabulonum, A. sabulosus, A. salmonis, A. saurinus, A. scaphoides, A. schmolliae, A. sclerocarpus, A. scopulorum, A. sepultipes, A. serenoi, A. sericoleucus, A. serpens, A. sesquiflorus, A. sheldonii, A. shevockii, A. shortianus, A. siliceus, A. simplicifolius, A. sinuatus, A. solitarius, A. sophoroides, A. soxmaniorum, A. spaldingii, A. sparsiflorus, A. spatulatus, A. speirocarpus, A. straturensis, A. striatiflorus, A. subcinereus, A. subvestitus, A. succumbens, A. tegetarioides, A. tener, A. tennesseensis, A. tephrodes, A. terminalis, A. tetrapterus, A. thurberi, A. tibetanus, A. tidestromii, A. tiehmii, A. titanophilus, A. toanus, A. toquimanus, A. tortipes, A. traskiae, A. tricarinatus, A. trichopodus, A. tridactylicus, A. troglodytus, A. tweedyi, A. tyghensis, A. umbellatus, A. umbraticus, A. uncialis, A. utahensis, A. vaccarum, A. vallaris, A. vexilliflexus, A. villosus, A. wardii, A. waterfallii, A. webberi, A. welshii, A. wetherillii, A. whitneyi, A. williamsii, A. wingatanus, A. wittmannii, A. woodruffii, A. wootonii, A. wrightii, A. xiphoides, A. yoderwilliamsii, A. zionis
Subordinate taxa
A. mollissimus var. bigelovii, A. mollissimus var. coryi, A. mollissimus var. earlei, A. mollissimus var. marcidus, A. mollissimus var. matthewsii, A. mollissimus var. mogollonicus, A. mollissimus var. mollissimus, A. mollissimus var. thompsoniae
Synonyms A. bigelovii
Name authority (A. Gray) Barneby: Mem. New York Bot. Gard. 13: 742. (1964) Torrey: Ann. Lyceum Nat.Hist. New York 2: 178. (1827)
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