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Missouri milk-vetch

Missouri milk-vetch

Habit Plants usually shortly caulescent, sometimes subacaulescent. Plants sometimes loosely tuft-forming, shortly caulescent, subacaulescent, or caulescent, 5–15(–20) cm, strigose or strigulose, hairs malpighian; from subterranean caudex.
Stems

to 15 cm.

prostrate and radiating, to 15(–20) cm, internodes often concealed by stipules, strigose or strigulose.

Leaves

(2–)4–14 cm;

stipules sometimes connate-sheathing at proximal nodes (vars. humistratus and missouriensis), (2–)3–9 mm, firm or submembranous becoming scarious or papery;

leaflets (5–)11–17(–21), blades elliptic to obovate, 3.5–13(–17) mm, apex acute, apiculate, or obtuse, surfaces strigose to strigulose.

Racemes

(3–)5–15-flowered.

(3–)4–15-flowered, flowers ascending to spreading;

axis (0.5–)1–4 cm in fruit;

bracts 2.5–8 mm;

bracteoles 0–2.

Peduncles

ascending, (1.5–)3.5–11 cm.

Pedicels

1–3.5 mm.

Flowers

calyx 9–12(–14.3) mm, tube 6.3–9(–9.3) mm, lobes 1.4–4.5(–5.3) mm;

corolla usually pink-purple, rarely white;

banner (14.5–)16–22(–24) mm;

keel (11.5–)12.8–17.3(–18.5) mm.

9.5–22(–24) mm;

calyx purple-tinged, cylindric or deeply campanulate, 5–13(–14.3) mm, strigulose, tube 4.1–10 mm, lobes subulate, (0.7–)1.4–4.5(–5.3) mm;

corolla usually pink-purple, purple, lavender, or violet-red, rarely white;

banner recurved through 45°;

keel 8.9–17.3(–18.5) mm.

Legumes

ascending, initially subterete or ± dorsiventrally compressed, ± straight, subsymmetrically oblong-ellipsoid, ± laterally compressed and obtuse-angled when mature, 15–28(–30) × (4–)5–9(–10) mm, subunilocular, base obtuse or sometimes cuneate, apex abruptly contracted into subulate, pungent beak, sutures prominent, strigulose.

usually persistent, ascending or descending, brownish becoming black, straight or slightly incurved, subsymmetrically or obliquely oblong, oblong-ellipsoid, or ellipsoid, subterete, or dorsiventrally or ± laterally compressed, (11–)14–28(–30) × (4–)5–9(–10) mm, unilocular or subunilocular, distinctly beaked, fleshy becoming stiffly leathery or subligneous, walls much less than 1 mm thick, strigulose, strigose, or glabrous.

Seeds

(33–)40–50(–56).

(33–)35–55(or 56).

2n

= 22.

= 22.

Astragalus missouriensis var. missouriensis

Astragalus missouriensis

Phenology Flowering late Mar–Jul.
Habitat Prairies, valleys, hillsides, dry open places, on limestone, shale, sandstone, or gyp­sum substrates.
Elevation 300–2400 m. (1000–7900 ft.)
Distribution
from FNA
AZ; CO; IA; KS; MN; MT; ND; NE; NM; OK; SD; TX; WY; AB; MB; SK
[BONAP county map]
from USDA
w North America
[WildflowerSearch map]
[BONAP county map]
Discussion

There is an irregular cline in flower size of var. missouriensis, from smallest in the north to largest in the south; caulescent forms appear to be more common southward (D. Isely 1998).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

From the central Canadian provinces to trans-Pecos Texas and western Iowa, Astragalus missouriensis is remarkably uniform morphologically. In the Four Corners region of the Southwest, it has undergone considerable differentiation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyces 5–5.8 mm; banners 9.5–11.8 mm; keel 8.9–10.6 mm; wc New Mexico, ne Arizona.
var. mimetes
1. Calyces 6–13(–14.3) mm; banners 14.5–22(–24) mm; keel (11.5–)12.8–17.3(–18.5) mm; Prairie Provinces southward to Arizona and Texas, eastward to Iowa and Minnesota.
→ 2
2. Legumes subsymmetrically oblong-ellipsoid, ± straight, initially subterete or ± dorsiven­trally compressed, at maturity ± laterally compressed and obtuse-angled; Prairie Provinces south­ward to Arizona and Texas, eastward to Iowa and Minnesota.
var. missouriensis
2. Legumes ellipsoid or oblong-ellipsoid, lunately incurved, dorsiventrally com­pressed, apices obcompressed proximal to incurved beak; Four Corners area.
→ 3
3. Plants subacaulescent to shortly cau­lescent; stems to 10 cm; racemes 4–8-flowered; Apache County, Arizona, Garfield to La Plata and Montezuma coun­ties, Colorado, San Juan County, New Mexico, Grand and San Juan coun­ties, Utah.
var. amphibolus
3. Plants caulescent; stems 10–15(–20) cm; racemes 9–12-flowered; Archuleta, Hins­dale, and La Plata counties, Colorado, adja­cent Rio Arriba County, New Mexico.
var. humistratus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Astragalus > sect. Argophylli > Astragalus missouriensis Fabaceae > subfam. Faboideae > Astragalus > sect. Argophylli
Sibling taxa
A. missouriensis var. amphibolus, A. missouriensis var. humistratus, A. missouriensis var. mimetes
A. accidens, A. accumbens, A. ackermanii, A. acutirostris, A. adanus, A. aequalis, A. agnicidus, A. agrestis, A. albens, A. albulus, A. allochrous, A. alpinus, A. altus, A. alvordensis, A. amblytropis, A. americanus, A. amnis-amissi, A. amphioxys, A. ampullarioides, A. ampullarius, A. andersonii, A. anisus, A. anserinus, A. applegatei, A. aquilonius, A. aretioides, A. argophyllus, A. aridus, A. arizonicus, A. arrectus, A. arthurii, A. asclepiadoides, A. asotinensis, A. asymmetricus, A. atratus, A. atropubescens, A. atwoodii, A. austiniae, A. australis, A. barnebyi, A. barrii, A. beathii, A. beatleyae, A. beckwithii, A. bernardinus, A. bibullatus, A. bicristatus, A. bisulcatus, A. bodinii, A. bolanderi, A. bourgovii, A. brandegeei, A. brauntonii, A. brazoensis, A. breweri, A. californicus, A. callithrix, A. calycosus, A. camptopus, A. canadensis, A. caricinus, A. casei, A. castaneiformis, A. castetteri, A. ceramicus, A. cerussatus, A. chamaeleuce, A. chamaemeniscus, A. chinensis, A. chloödes, A. chuskanus, A. cibarius, A. cicer, A. cimae, A. claranus, A. clevelandii, A. cliffordii, A. cobrensis, A. coccineus, A. collinus, A. coltonii, A. columbianus, A. concordius, A. congdonii, A. conjunctus, A. consobrinus, A. contortuplicatus, A. convallarius, A. cottamii, A. crassicarpus, A. cremnophylax, A. cronquistii, A. crotalariae, A. curtipes, A. curvicarpus, A. cusickii, A. cutleri, A. cyaneus, A. cymboides, A. deanei, A. debequaeus, A. desereticus, A. desperatus, A. deterior, A. detritalis, A. diaphanus, A. didymocarpus, A. distortus, A. diversifolius, A. douglasii, A. drabelliformis, A. drummondii, A. duchesnensis, A. eastwoodiae, A. egglestonii, A. emoryanus, A. endopterus, A. ensiformis, A. episcopus, A. equisolensis, A. eremiticus, A. ertterae, A. eucosmus, A. eurekensis, A. eurylobus, A. falcatus, A. feensis, A. filipes, A. flavus, A. flexuosus, A. fucatus, A. funereus, A. gambelianus, A. geyeri, A. gibbsii, A. giganteus, A. gilensis, A. gilmanii, A. gilviflorus, A. glycyphyllos, A. gracilis, A. grayi, A. gypsodes, A. hallii, A. hamiltonii, A. harrisonii, A. heilii, A. henrimontanensis, A. holmgreniorum, A. hoodianus, A. hornii, A. howellii, A. humillimus, A. humistratus, A. hyalinus, A. hypoxylus, A. inflexus, A. insularis, A. inversus, A. inyoensis, A. iodanthus, A. iodopetalus, A. iselyi, A. jaegerianus, A. jejunus, A. johannis-howellii, A. kelseyae, A. kentrophyta, A. kerrii, A. knightii, A. laccoliticus, A. lancearius, A. laxmannii, A. layneae, A. leibergii, A. lemmonii, A. lentiformis, A. lentiginosus, A. leptaleus, A. leptocarpus, A. leucolobus, A. limnocharis, A. lindheimeri, A. linifolius, A. loanus, A. lonchocarpus, A. lotiflorus, A. lutosus, A. lyallii, A. macrodon, A. magdalenae, A. malacoides, A. malacus, A. megacarpus, A. michauxii, A. microcymbus, A. microcystis, A. microlobium, A. micromerius, A. miguelensis, A. minthorniae, A. misellus, A. miser, A. moencoppensis, A. mohavensis, A. mollissimus, A. molybdenus, A. monoensis, A. montii, A. monumentalis, A. mulfordiae, A. multiflorus, A. musiniensis, A. naturitensis, A. neglectus, A. nelsonianus, A. neomexicanus, A. nevinii, A. newberryi, A. nidularius, A. nothoxys, A. nudisiliquus, A. nutans, A. nutriosensis, A. nuttallianus, A. nuttallii, A. nutzotinensis, A. nyensis, A. obcordatus, A. obscurus, A. oniciformis, A. oocalycis, A. oocarpus, A. oophorus, A. oreganus, A. osterhoutii, A. oxyphysus, A. pachypus, A. palmeri, A. panamintensis, A. pardalinus, A. parryi, A. pattersonii, A. pauperculus, A. paysonii, A. peckii, A. pectinatus, A. perianus, A. phoenix, A. pictiformis, A. pinonis, A. piscator, A. piutensis, A. plattensis, A. platytropis, A. polaris, A. pomonensis, A. porrectus, A. praelongus, A. preussii, A. proimanthus, A. proximus, A. pseudiodanthus, A. pterocarpus, A. pubentissimus, A. pulsiferae, A. puniceus, A. purshii, A. pycnostachyus, A. racemosus, A. rafaelensis, A. rattanii, A. ravenii, A. recurvus, A. reflexus, A. remotus, A. reventiformis, A. reventus, A. riparius, A. ripleyi, A. robbinsii, A. rusbyi, A. sabulonum, A. sabulosus, A. salmonis, A. saurinus, A. scaphoides, A. schmolliae, A. sclerocarpus, A. scopulorum, A. sepultipes, A. serenoi, A. sericoleucus, A. serpens, A. sesquiflorus, A. sheldonii, A. shevockii, A. shortianus, A. siliceus, A. simplicifolius, A. sinuatus, A. solitarius, A. sophoroides, A. soxmaniorum, A. spaldingii, A. sparsiflorus, A. spatulatus, A. speirocarpus, A. straturensis, A. striatiflorus, A. subcinereus, A. subvestitus, A. succumbens, A. tegetarioides, A. tener, A. tennesseensis, A. tephrodes, A. terminalis, A. tetrapterus, A. thurberi, A. tibetanus, A. tidestromii, A. tiehmii, A. titanophilus, A. toanus, A. toquimanus, A. tortipes, A. traskiae, A. tricarinatus, A. trichopodus, A. tridactylicus, A. troglodytus, A. tweedyi, A. tyghensis, A. umbellatus, A. umbraticus, A. uncialis, A. utahensis, A. vaccarum, A. vallaris, A. vexilliflexus, A. villosus, A. wardii, A. waterfallii, A. webberi, A. welshii, A. wetherillii, A. whitneyi, A. williamsii, A. wingatanus, A. wittmannii, A. woodruffii, A. wootonii, A. wrightii, A. xiphoides, A. yoderwilliamsii, A. zionis
Subordinate taxa
A. missouriensis var. amphibolus, A. missouriensis var. humistratus, A. missouriensis var. mimetes, A. missouriensis var. missouriensis
Name authority unknown Nuttall: Gen. N. Amer. Pl. 2: 99. (1818)
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