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archuleta milkvetch, Missouri milkvetch

Habit Plants caulescent. Herbs perennial (sometimes flowering as annual), usually tuft- or mat-forming, acaulescent, subacaulescent, or caulescent; caudex usually superficial or aerial, sometimes subterranean.
Stems

10–15(–20) cm.

(when present) obsolete, single, few, or several to many.

Leaves

odd-pinnate, usually petiolate, rarely short-petiolate or subsessile;

leaflets (1 or)3–39(–43).

Racemes

9–12-flowered.

subcapitate to loosely flowered, flowers erect, ascending, spreading, declined, or nodding, secund, and retrorsely imbricate.

Flowers

calyx 7.8–10 mm, tube 6–10 mm, lobes 1.5–3 mm;

corolla lavender, purple, or almost white, wing tips often white;

banner (17–)19–20.5 mm.

Corollas

purple, pink-purple, magenta-purple, violet, bluish, lilac, scarlet, ochroleucous, greenish white, or white, banner barely recurved (A. phoenix) or recurved through 20–50° (90–100° in A. accumbens), keel apex obtuse.

Calyx

tubes cylindric or deeply campanulate.

Legumes

ascending to descending, dorsiventrally compressed, lunately incurved, oblong-ellipsoid, (12–)17–20 × 6–9 mm, unilocular, apex obcompressed proximal to incurved beak, glabrous or sparsely strigulose.

usually deciduous, usually sessile, rarely subsessile or substipitate, gynophore sometimes present, usually ascending (humistrate), less often spreading or pendulous, subglobose to ellipsoid, narrowly lanceoloid, ovoid or oblong-ellipsoid, or lanceoloid-ovoid, straight or usually incurved, usually compressed dorsiventrally, sometimes obcompressed, 3-sided, turgid, or inflated, unilocular, subunilocular, or bilocular.

Seeds

33–40.

11–70.

Hairs

basifixed or malpighian.

Stipules

distinct (except anomalous forms of A. missouriensis, A. tephrodes, and A. zionis).

Astragalus missouriensis var. humistratus

Astragalus sect. Argophylli

Phenology Flowering May–Jul.
Habitat Oak brush with scattered pon­derosa pine on clay knolls, pinyon-juniper woodlands, associated with Lewis and Mancos formations.
Elevation 2100–2500 m. (6900–8200 ft.)
Distribution
from FNA
CO; NM
[BONAP county map]
w North America; n Mexico
Discussion

Variety humistratus is locally distributed in Archuleta, Hinsdale, and La Plata counties, Colorado, and adjacent Rio Arriba County, New Mexico. It is anomalous in its strongly caulescent but mat-forming habit, and the slightly or plainly connate stipules. R. C. Barneby (1964) suggested a hybrid origin between Astragalus missouriensis and A. humistratus.

Variety humistratus is in the Center for Plant Conservation’s National Collection of Endangered Plants.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 44 (44 in the flora).

Section Argophylli comprises ten subsections, widespread in western North America from southern British Columbia and Saskatchewan southward to northern Baja California, northern Sonora, and western Texas.

The subsections are: subsect. Argophylli (A. Gray) M. E. Jones (Astragalus argophyllus, A. callithrix, A. columbianus, A. cyaneus, A. desereticus, A. eurylobus, A. henrimontanensis, A. iodopetalus, A. piutensis, A. shortianus, A. tephrodes, A. tidestromii, A. uncialis, A. zionis); subsect. Pseudargophylli Barneby (A. feensis, A. waterfallii); subsect. Neomexicani Barneby (A. neomexicanus); subsect. Newberryani M. E. Jones (A. eurekensis, A. loanus, A. musiniensis, A. newberryi, A. phoenix, A. welshii); subsect. Concordi S. L. Welsh (A. concordius); subsect. Coccinei M. E. Jones (A. coccineus); subsect. Eriocarpi (A. Gray) Barneby (A. anserinus, A. funereus, A. inflexus, A. leucolobus, A. nudisiliquus, A. purshii, A. subvestitus, A. utahensis); subsect. Parryani Barneby (A. parryi); subsect. Missourienses M. E. Jones (A. accumbens, A. amphioxys, A. castaneiformis, A. chamaeleuce, A. consobrinus, A. cymboides, A. laccoliticus, A. missouriensis, A. piscator); and subsect. Anisi Barneby (A. anisus).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Astragalus > sect. Argophylli > Astragalus missouriensis Fabaceae > subfam. Faboideae > Astragalus
Sibling taxa
A. missouriensis var. amphibolus, A. missouriensis var. mimetes, A. missouriensis var. missouriensis
Subordinate taxa
Name authority Isely: Syst. Bot. 8: 423. (1983) A. Gray: Proc. Amer. Acad. Arts 6: 209. (1864)
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