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Mancos milkvetch, Missouri milkvetch

Missouri milk-vetch

Habit Plants subacaulescent to shortly caulescent. Plants sometimes loosely tuft-forming, shortly caulescent, subacaulescent, or caulescent, 5–15(–20) cm, strigose or strigulose, hairs malpighian; from subterranean caudex.
Stems

to 10 cm.

prostrate and radiating, to 15(–20) cm, internodes often concealed by stipules, strigose or strigulose.

Leaves

(2–)4–14 cm;

stipules sometimes connate-sheathing at proximal nodes (vars. humistratus and missouriensis), (2–)3–9 mm, firm or submembranous becoming scarious or papery;

leaflets (5–)11–17(–21), blades elliptic to obovate, 3.5–13(–17) mm, apex acute, apiculate, or obtuse, surfaces strigose to strigulose.

Racemes

4–8-flowered.

(3–)4–15-flowered, flowers ascending to spreading;

axis (0.5–)1–4 cm in fruit;

bracts 2.5–8 mm;

bracteoles 0–2.

Peduncles

ascending, (1.5–)3.5–11 cm.

Pedicels

1–3.5 mm.

Flowers

calyx 8.5–13 mm, tube 7–10 mm, lobes 1.5–3 mm;

corolla usually pink-purple, rarely white;

banner (14.5–)16–22(–24) mm;

keel (11.5–)12.8–17.3(–18.5) mm.

9.5–22(–24) mm;

calyx purple-tinged, cylindric or deeply campanulate, 5–13(–14.3) mm, strigulose, tube 4.1–10 mm, lobes subulate, (0.7–)1.4–4.5(–5.3) mm;

corolla usually pink-purple, purple, lavender, or violet-red, rarely white;

banner recurved through 45°;

keel 8.9–17.3(–18.5) mm.

Legumes

sometimes deciduous, ascending to descending, dorsiventrally compressed, lunately incurved, ellipsoid, (11–)15–25 × 7–9 mm, unilocular, apex obcompressed proximal to incurved beak, strigose.

usually persistent, ascending or descending, brownish becoming black, straight or slightly incurved, subsymmetrically or obliquely oblong, oblong-ellipsoid, or ellipsoid, subterete, or dorsiventrally or ± laterally compressed, (11–)14–28(–30) × (4–)5–9(–10) mm, unilocular or subunilocular, distinctly beaked, fleshy becoming stiffly leathery or subligneous, walls much less than 1 mm thick, strigulose, strigose, or glabrous.

Seeds

35–55.

(33–)35–55(or 56).

2n

= 22.

Astragalus missouriensis var. amphibolus

Astragalus missouriensis

Phenology Flowering May–Jul.
Habitat Pinyon-juniper and sagebrush communities, on igneous or sandstone outcrops or sub­strates.
Elevation 1600–2500 m. (5200–8200 ft.)
Distribution
from FNA
AZ; CO; NM; UT
[BONAP county map]
from USDA
w North America
[WildflowerSearch map]
[BONAP county map]
Discussion

The fruits of var. amphibolus are initially dorsiventrally compressed, and ultimately dehisce apically while still attached to the inflorescence (though sometimes deciduous). The fruits have a lateral ridge down each valve, with the valves separated by more or less prominent bicarinate keels. In these features, along with the typically persistent fruits, the plants can be distinguished from the similar Astragalus amphioxys var. amphioxys where their ranges are contiguous, as in northwestern New Mexico and vicinity. R. C. Barneby (1947b, 1964) suggested that hybridization occurs between the two taxa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

From the central Canadian provinces to trans-Pecos Texas and western Iowa, Astragalus missouriensis is remarkably uniform morphologically. In the Four Corners region of the Southwest, it has undergone considerable differentiation.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Calyces 5–5.8 mm; banners 9.5–11.8 mm; keel 8.9–10.6 mm; wc New Mexico, ne Arizona.
var. mimetes
1. Calyces 6–13(–14.3) mm; banners 14.5–22(–24) mm; keel (11.5–)12.8–17.3(–18.5) mm; Prairie Provinces southward to Arizona and Texas, eastward to Iowa and Minnesota.
→ 2
2. Legumes subsymmetrically oblong-ellipsoid, ± straight, initially subterete or ± dorsiven­trally compressed, at maturity ± laterally compressed and obtuse-angled; Prairie Provinces south­ward to Arizona and Texas, eastward to Iowa and Minnesota.
var. missouriensis
2. Legumes ellipsoid or oblong-ellipsoid, lunately incurved, dorsiventrally com­pressed, apices obcompressed proximal to incurved beak; Four Corners area.
→ 3
3. Plants subacaulescent to shortly cau­lescent; stems to 10 cm; racemes 4–8-flowered; Apache County, Arizona, Garfield to La Plata and Montezuma coun­ties, Colorado, San Juan County, New Mexico, Grand and San Juan coun­ties, Utah.
var. amphibolus
3. Plants caulescent; stems 10–15(–20) cm; racemes 9–12-flowered; Archuleta, Hins­dale, and La Plata counties, Colorado, adja­cent Rio Arriba County, New Mexico.
var. humistratus
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Astragalus > sect. Argophylli > Astragalus missouriensis Fabaceae > subfam. Faboideae > Astragalus > sect. Argophylli
Sibling taxa
A. missouriensis var. humistratus, A. missouriensis var. mimetes, A. missouriensis var. missouriensis
A. accidens, A. accumbens, A. ackermanii, A. acutirostris, A. adanus, A. aequalis, A. agnicidus, A. agrestis, A. albens, A. albulus, A. allochrous, A. alpinus, A. altus, A. alvordensis, A. amblytropis, A. americanus, A. amnis-amissi, A. amphioxys, A. ampullarioides, A. ampullarius, A. andersonii, A. anisus, A. anserinus, A. applegatei, A. aquilonius, A. aretioides, A. argophyllus, A. aridus, A. arizonicus, A. arrectus, A. arthurii, A. asclepiadoides, A. asotinensis, A. asymmetricus, A. atratus, A. atropubescens, A. atwoodii, A. austiniae, A. australis, A. barnebyi, A. barrii, A. beathii, A. beatleyae, A. beckwithii, A. bernardinus, A. bibullatus, A. bicristatus, A. bisulcatus, A. bodinii, A. bolanderi, A. bourgovii, A. brandegeei, A. brauntonii, A. brazoensis, A. breweri, A. californicus, A. callithrix, A. calycosus, A. camptopus, A. canadensis, A. caricinus, A. casei, A. castaneiformis, A. castetteri, A. ceramicus, A. cerussatus, A. chamaeleuce, A. chamaemeniscus, A. chinensis, A. chloödes, A. chuskanus, A. cibarius, A. cicer, A. cimae, A. claranus, A. clevelandii, A. cliffordii, A. cobrensis, A. coccineus, A. collinus, A. coltonii, A. columbianus, A. concordius, A. congdonii, A. conjunctus, A. consobrinus, A. contortuplicatus, A. convallarius, A. cottamii, A. crassicarpus, A. cremnophylax, A. cronquistii, A. crotalariae, A. curtipes, A. curvicarpus, A. cusickii, A. cutleri, A. cyaneus, A. cymboides, A. deanei, A. debequaeus, A. desereticus, A. desperatus, A. deterior, A. detritalis, A. diaphanus, A. didymocarpus, A. distortus, A. diversifolius, A. douglasii, A. drabelliformis, A. drummondii, A. duchesnensis, A. eastwoodiae, A. egglestonii, A. emoryanus, A. endopterus, A. ensiformis, A. episcopus, A. equisolensis, A. eremiticus, A. ertterae, A. eucosmus, A. eurekensis, A. eurylobus, A. falcatus, A. feensis, A. filipes, A. flavus, A. flexuosus, A. fucatus, A. funereus, A. gambelianus, A. geyeri, A. gibbsii, A. giganteus, A. gilensis, A. gilmanii, A. gilviflorus, A. glycyphyllos, A. gracilis, A. grayi, A. gypsodes, A. hallii, A. hamiltonii, A. harrisonii, A. heilii, A. henrimontanensis, A. holmgreniorum, A. hoodianus, A. hornii, A. howellii, A. humillimus, A. humistratus, A. hyalinus, A. hypoxylus, A. inflexus, A. insularis, A. inversus, A. inyoensis, A. iodanthus, A. iodopetalus, A. iselyi, A. jaegerianus, A. jejunus, A. johannis-howellii, A. kelseyae, A. kentrophyta, A. kerrii, A. knightii, A. laccoliticus, A. lancearius, A. laxmannii, A. layneae, A. leibergii, A. lemmonii, A. lentiformis, A. lentiginosus, A. leptaleus, A. leptocarpus, A. leucolobus, A. limnocharis, A. lindheimeri, A. linifolius, A. loanus, A. lonchocarpus, A. lotiflorus, A. lutosus, A. lyallii, A. macrodon, A. magdalenae, A. malacoides, A. malacus, A. megacarpus, A. michauxii, A. microcymbus, A. microcystis, A. microlobium, A. micromerius, A. miguelensis, A. minthorniae, A. misellus, A. miser, A. moencoppensis, A. mohavensis, A. mollissimus, A. molybdenus, A. monoensis, A. montii, A. monumentalis, A. mulfordiae, A. multiflorus, A. musiniensis, A. naturitensis, A. neglectus, A. nelsonianus, A. neomexicanus, A. nevinii, A. newberryi, A. nidularius, A. nothoxys, A. nudisiliquus, A. nutans, A. nutriosensis, A. nuttallianus, A. nuttallii, A. nutzotinensis, A. nyensis, A. obcordatus, A. obscurus, A. oniciformis, A. oocalycis, A. oocarpus, A. oophorus, A. oreganus, A. osterhoutii, A. oxyphysus, A. pachypus, A. palmeri, A. panamintensis, A. pardalinus, A. parryi, A. pattersonii, A. pauperculus, A. paysonii, A. peckii, A. pectinatus, A. perianus, A. phoenix, A. pictiformis, A. pinonis, A. piscator, A. piutensis, A. plattensis, A. platytropis, A. polaris, A. pomonensis, A. porrectus, A. praelongus, A. preussii, A. proimanthus, A. proximus, A. pseudiodanthus, A. pterocarpus, A. pubentissimus, A. pulsiferae, A. puniceus, A. purshii, A. pycnostachyus, A. racemosus, A. rafaelensis, A. rattanii, A. ravenii, A. recurvus, A. reflexus, A. remotus, A. reventiformis, A. reventus, A. riparius, A. ripleyi, A. robbinsii, A. rusbyi, A. sabulonum, A. sabulosus, A. salmonis, A. saurinus, A. scaphoides, A. schmolliae, A. sclerocarpus, A. scopulorum, A. sepultipes, A. serenoi, A. sericoleucus, A. serpens, A. sesquiflorus, A. sheldonii, A. shevockii, A. shortianus, A. siliceus, A. simplicifolius, A. sinuatus, A. solitarius, A. sophoroides, A. soxmaniorum, A. spaldingii, A. sparsiflorus, A. spatulatus, A. speirocarpus, A. straturensis, A. striatiflorus, A. subcinereus, A. subvestitus, A. succumbens, A. tegetarioides, A. tener, A. tennesseensis, A. tephrodes, A. terminalis, A. tetrapterus, A. thurberi, A. tibetanus, A. tidestromii, A. tiehmii, A. titanophilus, A. toanus, A. toquimanus, A. tortipes, A. traskiae, A. tricarinatus, A. trichopodus, A. tridactylicus, A. troglodytus, A. tweedyi, A. tyghensis, A. umbellatus, A. umbraticus, A. uncialis, A. utahensis, A. vaccarum, A. vallaris, A. vexilliflexus, A. villosus, A. wardii, A. waterfallii, A. webberi, A. welshii, A. wetherillii, A. whitneyi, A. williamsii, A. wingatanus, A. wittmannii, A. woodruffii, A. wootonii, A. wrightii, A. xiphoides, A. yoderwilliamsii, A. zionis
Subordinate taxa
A. missouriensis var. amphibolus, A. missouriensis var. humistratus, A. missouriensis var. mimetes, A. missouriensis var. missouriensis
Name authority Barneby: Amer. Midl. Naturalist 37: 447. (1947) Nuttall: Gen. N. Amer. Pl. 2: 99. (1818)
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