Mancos milkvetch, Missouri milkvetch
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Missouri milk-vetch
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Plants subacaulescent to shortly caulescent. |
Plants sometimes loosely tuft-forming, shortly caulescent, subacaulescent, or caulescent, 5–15(–20) cm, strigose or strigulose, hairs malpighian; from subterranean caudex. |
to 10 cm. |
prostrate and radiating, to 15(–20) cm, internodes often concealed by stipules, strigose or strigulose. |
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(2–)4–14 cm; stipules sometimes connate-sheathing at proximal nodes (vars. humistratus and missouriensis), (2–)3–9 mm, firm or submembranous becoming scarious or papery; leaflets (5–)11–17(–21), blades elliptic to obovate, 3.5–13(–17) mm, apex acute, apiculate, or obtuse, surfaces strigose to strigulose. |
4–8-flowered. |
(3–)4–15-flowered, flowers ascending to spreading; axis (0.5–)1–4 cm in fruit; bracts 2.5–8 mm; bracteoles 0–2. |
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ascending, (1.5–)3.5–11 cm. |
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1–3.5 mm. |
calyx 8.5–13 mm, tube 7–10 mm, lobes 1.5–3 mm; corolla usually pink-purple, rarely white; banner (14.5–)16–22(–24) mm; keel (11.5–)12.8–17.3(–18.5) mm. |
9.5–22(–24) mm; calyx purple-tinged, cylindric or deeply campanulate, 5–13(–14.3) mm, strigulose, tube 4.1–10 mm, lobes subulate, (0.7–)1.4–4.5(–5.3) mm; corolla usually pink-purple, purple, lavender, or violet-red, rarely white; banner recurved through 45°; keel 8.9–17.3(–18.5) mm. |
sometimes deciduous, ascending to descending, dorsiventrally compressed, lunately incurved, ellipsoid, (11–)15–25 × 7–9 mm, unilocular, apex obcompressed proximal to incurved beak, strigose. |
usually persistent, ascending or descending, brownish becoming black, straight or slightly incurved, subsymmetrically or obliquely oblong, oblong-ellipsoid, or ellipsoid, subterete, or dorsiventrally or ± laterally compressed, (11–)14–28(–30) × (4–)5–9(–10) mm, unilocular or subunilocular, distinctly beaked, fleshy becoming stiffly leathery or subligneous, walls much less than 1 mm thick, strigulose, strigose, or glabrous. |
35–55. |
(33–)35–55(or 56). |
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= 22. |
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Flowering May–Jul. |
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Pinyon-juniper and sagebrush communities, on igneous or sandstone outcrops or substrates. |
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1600–2500 m. (5200–8200 ft.) |
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AZ; CO; NM; UT |
w North America
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The fruits of var. amphibolus are initially dorsiventrally compressed, and ultimately dehisce apically while still attached to the inflorescence (though sometimes deciduous). The fruits have a lateral ridge down each valve, with the valves separated by more or less prominent bicarinate keels. In these features, along with the typically persistent fruits, the plants can be distinguished from the similar Astragalus amphioxys var. amphioxys where their ranges are contiguous, as in northwestern New Mexico and vicinity. R. C. Barneby (1947b, 1964) suggested that hybridization occurs between the two taxa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 4 (4 in the flora). From the central Canadian provinces to trans-Pecos Texas and western Iowa, Astragalus missouriensis is remarkably uniform morphologically. In the Four Corners region of the Southwest, it has undergone considerable differentiation. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Calyces 5–5.8 mm; banners 9.5–11.8 mm; keel 8.9–10.6 mm; wc New Mexico, ne Arizona. | var. mimetes |
1. Calyces 6–13(–14.3) mm; banners 14.5–22(–24) mm; keel (11.5–)12.8–17.3(–18.5) mm; Prairie Provinces southward to Arizona and Texas, eastward to Iowa and Minnesota. | → 2 |
2. Legumes subsymmetrically oblong-ellipsoid, ± straight, initially subterete or ± dorsiventrally compressed, at maturity ± laterally compressed and obtuse-angled; Prairie Provinces southward to Arizona and Texas, eastward to Iowa and Minnesota. | var. missouriensis |
2. Legumes ellipsoid or oblong-ellipsoid, lunately incurved, dorsiventrally compressed, apices obcompressed proximal to incurved beak; Four Corners area. | → 3 |
3. Plants subacaulescent to shortly caulescent; stems to 10 cm; racemes 4–8-flowered; Apache County, Arizona, Garfield to La Plata and Montezuma counties, Colorado, San Juan County, New Mexico, Grand and San Juan counties, Utah. | var. amphibolus |
3. Plants caulescent; stems 10–15(–20) cm; racemes 9–12-flowered; Archuleta, Hinsdale, and La Plata counties, Colorado, adjacent Rio Arriba County, New Mexico. | var. humistratus |
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FNA vol. 11. |
FNA vol. 11. |
Fabaceae > subfam. Faboideae > Astragalus > sect. Argophylli > Astragalus missouriensis |
Fabaceae > subfam. Faboideae > Astragalus > sect. Argophylli |
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A. accidens, A. accumbens, A. ackermanii, A. acutirostris, A. adanus, A. aequalis, A. agnicidus, A. agrestis, A. albens, A. albulus, A. allochrous, A. alpinus, A. altus, A. alvordensis, A. amblytropis, A. americanus, A. amnis-amissi, A. amphioxys, A. ampullarioides, A. ampullarius, A. andersonii, A. anisus, A. anserinus, A. applegatei, A. aquilonius, A. aretioides, A. argophyllus, A. aridus, A. arizonicus, A. arrectus, A. arthurii, A. asclepiadoides, A. asotinensis, A. asymmetricus, A. atratus, A. atropubescens, A. atwoodii, A. austiniae, A. australis, A. barnebyi, A. barrii, A. beathii, A. beatleyae, A. beckwithii, A. bernardinus, A. bibullatus, A. bicristatus, A. bisulcatus, A. bodinii, A. bolanderi, A. bourgovii, A. brandegeei, A. brauntonii, A. brazoensis, A. breweri, A. californicus, A. callithrix, A. calycosus, A. camptopus, A. canadensis, A. caricinus, A. casei, A. castaneiformis, A. castetteri, A. ceramicus, A. cerussatus, A. chamaeleuce, A. chamaemeniscus, A. chinensis, A. chloödes, A. chuskanus, A. cibarius, A. cicer, A. cimae, A. claranus, A. clevelandii, A. cliffordii, A. cobrensis, A. coccineus, A. collinus, A. coltonii, A. columbianus, A. concordius, A. congdonii, A. conjunctus, A. consobrinus, A. contortuplicatus, A. convallarius, A. cottamii, A. crassicarpus, A. cremnophylax, A. cronquistii, A. crotalariae, A. curtipes, A. curvicarpus, A. cusickii, A. cutleri, A. cyaneus, A. cymboides, A. deanei, A. debequaeus, A. desereticus, A. desperatus, A. deterior, A. detritalis, A. diaphanus, A. didymocarpus, A. distortus, A. diversifolius, A. douglasii, A. drabelliformis, A. drummondii, A. duchesnensis, A. eastwoodiae, A. egglestonii, A. emoryanus, A. endopterus, A. ensiformis, A. episcopus, A. equisolensis, A. eremiticus, A. ertterae, A. eucosmus, A. eurekensis, A. eurylobus, A. falcatus, A. feensis, A. filipes, A. flavus, A. flexuosus, A. fucatus, A. funereus, A. gambelianus, A. geyeri, A. gibbsii, A. giganteus, A. gilensis, A. gilmanii, A. gilviflorus, A. glycyphyllos, A. gracilis, A. grayi, A. gypsodes, A. hallii, A. hamiltonii, A. harrisonii, A. heilii, A. henrimontanensis, A. holmgreniorum, A. hoodianus, A. hornii, A. howellii, A. humillimus, A. humistratus, A. hyalinus, A. hypoxylus, A. inflexus, A. insularis, A. inversus, A. inyoensis, A. iodanthus, A. iodopetalus, A. iselyi, A. jaegerianus, A. jejunus, A. johannis-howellii, A. kelseyae, A. kentrophyta, A. kerrii, A. knightii, A. laccoliticus, A. lancearius, A. laxmannii, A. layneae, A. leibergii, A. lemmonii, A. lentiformis, A. lentiginosus, A. leptaleus, A. leptocarpus, A. leucolobus, A. limnocharis, A. lindheimeri, A. linifolius, A. loanus, A. lonchocarpus, A. lotiflorus, A. lutosus, A. lyallii, A. macrodon, A. magdalenae, A. malacoides, A. malacus, A. megacarpus, A. michauxii, A. microcymbus, A. microcystis, A. microlobium, A. micromerius, A. miguelensis, A. minthorniae, A. misellus, A. miser, A. moencoppensis, A. mohavensis, A. mollissimus, A. molybdenus, A. monoensis, A. montii, A. monumentalis, A. mulfordiae, A. multiflorus, A. musiniensis, A. naturitensis, A. neglectus, A. nelsonianus, A. neomexicanus, A. nevinii, A. newberryi, A. nidularius, A. nothoxys, A. nudisiliquus, A. nutans, A. nutriosensis, A. nuttallianus, A. nuttallii, A. nutzotinensis, A. nyensis, A. obcordatus, A. obscurus, A. oniciformis, A. oocalycis, A. oocarpus, A. oophorus, A. oreganus, A. osterhoutii, A. oxyphysus, A. pachypus, A. palmeri, A. panamintensis, A. pardalinus, A. parryi, A. pattersonii, A. pauperculus, A. paysonii, A. peckii, A. pectinatus, A. perianus, A. phoenix, A. pictiformis, A. pinonis, A. piscator, A. piutensis, A. plattensis, A. platytropis, A. polaris, A. pomonensis, A. porrectus, A. praelongus, A. preussii, A. proimanthus, A. proximus, A. pseudiodanthus, A. pterocarpus, A. pubentissimus, A. pulsiferae, A. puniceus, A. purshii, A. pycnostachyus, A. racemosus, A. rafaelensis, A. rattanii, A. ravenii, A. recurvus, A. reflexus, A. remotus, A. reventiformis, A. reventus, A. riparius, A. ripleyi, A. robbinsii, A. rusbyi, A. sabulonum, A. sabulosus, A. salmonis, A. saurinus, A. scaphoides, A. schmolliae, A. sclerocarpus, A. scopulorum, A. sepultipes, A. serenoi, A. sericoleucus, A. serpens, A. sesquiflorus, A. sheldonii, A. shevockii, A. shortianus, A. siliceus, A. simplicifolius, A. sinuatus, A. solitarius, A. sophoroides, A. soxmaniorum, A. spaldingii, A. sparsiflorus, A. spatulatus, A. speirocarpus, A. straturensis, A. striatiflorus, A. subcinereus, A. subvestitus, A. succumbens, A. tegetarioides, A. tener, A. tennesseensis, A. tephrodes, A. terminalis, A. tetrapterus, A. thurberi, A. tibetanus, A. tidestromii, A. tiehmii, A. titanophilus, A. toanus, A. toquimanus, A. tortipes, A. traskiae, A. tricarinatus, A. trichopodus, A. tridactylicus, A. troglodytus, A. tweedyi, A. tyghensis, A. umbellatus, A. umbraticus, A. uncialis, A. utahensis, A. vaccarum, A. vallaris, A. vexilliflexus, A. villosus, A. wardii, A. waterfallii, A. webberi, A. welshii, A. wetherillii, A. whitneyi, A. williamsii, A. wingatanus, A. wittmannii, A. woodruffii, A. wootonii, A. wrightii, A. xiphoides, A. yoderwilliamsii, A. zionis |
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Barneby: Amer. Midl. Naturalist 37: 447. (1947) |
Nuttall: Gen. N. Amer. Pl. 2: 99. (1818) |
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