Douglas kentrophyta, Douglas' spiny milkvetch, spiny millk-vetch, thistle milk-vetch
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kentrophyta, Nuttall's kentrophyta, spiny milk-vetch, thistle milk-vetch
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Plants prostrate, densely mat-forming, becoming suffruticose. |
Plants often mat- or cushion-forming, sometimes suffruticose basally, 15–45 cm, usually strigulose, sometimes villosulous or villous, hairs basifixed or malpighian; from caudex bearing stolonlike, creeping stems. |
and herbage strigulose, hairs basifixed. |
prostrate, decumbent, spreading, or erect, compact to elongate, usually strigulose, sometimes villosulous or villous. |
1–1.7 cm; stipules dimorphic, 2–5 mm; leaflets 5(or 7), blades 5–12 mm, surfaces pubescent. |
(0.2–)0.4–2.6 cm; stipules connate-sheathing, sometimes distinct at distal nodes, 1–12 mm, scarious throughout, or herbaceous becoming papery, or stiff and spiny, apex acuminate, spinulose, or callous-mucronate; leaflets (3 or)5–9, blades decurrent, continuous with rachis, linear to narrowly elliptic or lanceolate, (1–)3–15(–17) mm, apex spinulose, surfaces usually strigose, rarely glabrous adaxially. |
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1–3-flowered, flowers declined; axis to 0.5 cm in fruit; bracts 0.8–3.5 mm; bracteoles 0. |
subobsolete. |
subobsolete or 0.1–0.6(–3) cm. |
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0.5–2 mm. |
5.8 mm; calyx 4.7–5.2 mm, tube 2.2–2.4 mm, lobes subulate, spinulose, 2.3–3 mm; corolla whitish. |
(3.9–)4–10 mm; calyx (2–)2.4–8.3 mm, strigose, tube 1.2–2.8(–3.3) mm, lobes subulate, setaceous, or spinulose, (0.5–)1.4–5 mm; corolla usually pink-purple or whitish, sometimes pink-, lilac-, or purple-tinged; banner recurved through 45°; keel 3–6 mm. |
lanceoloid, 5–5.5 × 2 mm. |
declined or spreading, stramineous, straight or slightly curved, ellipsoid to oblong-ellipsoid, lenticular, ovoid, ovoid-lenticular, lanceoloid-ellipsoid, or lanceoloid-acuminate, laterally compressed, 3–10 × (0.6–)1.4–4 mm, papery, strigose; sessile. |
2. |
2–8. |
= 24. |
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Flowering Jun. |
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Sandy substrates. |
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150–400 m. (500–1300 ft.) |
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OR; WA |
w North America
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Variety douglasii is known from only a few, vague historical records, probably taken near the present city of Walla Walla near the Great Bend of the Columbia River, and appears to be extinct. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 9 (9 in the flora). The varieties of Astragalus kentrophyta are distinguished from other astragali by the few leaflets continuous with the leafstalk while blades are simultaneously apically mucronate or spinulose, and by their racemes, usually subsessile in leaf axils, bearing one to three small flowers. There are two major groups: alpine or upper elevation, prostrate-matted to pulvinate-depressed plants, usually with bright purple flowers and basifixed hairs, with five to eight seeds; and lower elevation forms that range from prostrate to bushy and ascending, with white or lavender flowers, hairs sometimes malpighian, and two to five ovules. With the exception of the widespread var. tegetarius, the varieties are largely allopatric. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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1. Hairs basifixed; seeds (3–)5–8; corollas purple, purplish, or whitish and keel purple or pink- or purplish-tipped. | → 2 |
2. Leaflets (3–)5–9, blades 1–9 mm; plants forming cushions 5–40(–50) cm wide; widespread, extending into California only to Inyo and White mountains and c High Sierra Nevada. | var. tegetarius |
2. Leaflets 3(–7), blades 3–7 mm; plants forming cushions to 15 cm wide; sc Sierra Nevada, California. | var. danaus |
1. Hairs malpighian or basifixed; seeds 2–4(–8); corollas whitish, keel apices sometimes pink or lilac-tinged or purplish (except pink-purple in var. coloradoensis, rarely in var. elatus). | → 3 |
3. Calyces 6–8.3 mm, lobes 3.4–5 mm; legumes (5–)7–10 × 2.8–4 mm; seeds 4–8; canyons of Colorado River in nc Arizona and sc Utah. | var. coloradoensis |
3. Calyces 2.4–5.2 mm, lobes 0.7–3 mm; legumes 3–7(–7.5) × 1.4–3.8 mm; seeds 2–4; California to Alberta, eastward to New Mexico and South Dakota. | → 4 |
4. Legumes ovoid-lenticular, ovoid, or lenticular. | → 5 |
5. Stipules uniform, connate to 1/2 their lengths, spinulose; calyx lobes 0.7–1.5 mm; n New Mexico. | var. neomexicanus |
5. Stipules dimorphic, those at most proximal nodes connate into short, bidentate sheath, those at distal nodes longer, connate at base only, lanceolate or lanceolate-acuminate, often without spines; calyx lobes 1.4–3 mm; North Dakota west to Alberta, southward to Colorado and Oregon. | → 6 |
6. Leaflet blades glabrous or glabrescent adaxially, pubescent abaxially, hairs basifixed or obscurely malpighian; legumes (3.5–)4–7 × 2.4–3.8 mm; s Wyoming east of Continental Divide to Alberta and Saskatchewan, southward to Colorado, South Dakota, and w Nebraska. | var. kentrophyta |
6. Leaflet blades pubescent, hairs malpighian; legumes 3–4.5 × 1.4–2 mm; sw Wyoming, ne Utah to se Oregon. | var. jessiae |
4. Legumes ovoid-acuminate, lanceoloid, or lanceoloid-acuminate. | → 7 |
7. Plants usually erect or assurgent, rarely trailing, often bushy, sometimes mat-forming; Arizona, California, Colorado, Nevada, New Mexico, Utah, Wyoming. | var. elatus |
7. Plants prostrate, mat- or cushion-forming; California, e, c Nevada, Oregon, se Washington. | → 8 |
8. Hairs basifixed; calyx lobes 2.3–3 mm; Oregon, se Washington. | var. douglasii |
8. Hairs malpighian; calyx lobes 1.8–2.6 mm; California, Nevada. | var. ungulatus |
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FNA vol. 11. |
FNA vol. 11. |
Fabaceae > subfam. Faboideae > Astragalus > sect. Ervoidei > Astragalus kentrophyta |
Fabaceae > subfam. Faboideae > Astragalus > sect. Ervoidei |
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A. accidens, A. accumbens, A. ackermanii, A. acutirostris, A. adanus, A. aequalis, A. agnicidus, A. agrestis, A. albens, A. albulus, A. allochrous, A. alpinus, A. altus, A. alvordensis, A. amblytropis, A. americanus, A. amnis-amissi, A. amphioxys, A. ampullarioides, A. ampullarius, A. andersonii, A. anisus, A. anserinus, A. applegatei, A. aquilonius, A. aretioides, A. argophyllus, A. aridus, A. arizonicus, A. arrectus, A. arthurii, A. asclepiadoides, A. asotinensis, A. asymmetricus, A. atratus, A. atropubescens, A. atwoodii, A. austiniae, A. australis, A. barnebyi, A. barrii, A. beathii, A. beatleyae, A. beckwithii, A. bernardinus, A. bibullatus, A. bicristatus, A. bisulcatus, A. bodinii, A. bolanderi, A. bourgovii, A. brandegeei, A. brauntonii, A. brazoensis, A. breweri, A. californicus, A. callithrix, A. calycosus, A. camptopus, A. canadensis, A. caricinus, A. casei, A. castaneiformis, A. castetteri, A. ceramicus, A. cerussatus, A. chamaeleuce, A. chamaemeniscus, A. chinensis, A. chloödes, A. chuskanus, A. cibarius, A. cicer, A. cimae, A. claranus, A. clevelandii, A. cliffordii, A. cobrensis, A. coccineus, A. collinus, A. coltonii, A. columbianus, A. concordius, A. congdonii, A. conjunctus, A. consobrinus, A. contortuplicatus, A. convallarius, A. cottamii, A. crassicarpus, A. cremnophylax, A. cronquistii, A. crotalariae, A. curtipes, A. curvicarpus, A. cusickii, A. cutleri, A. cyaneus, A. cymboides, A. deanei, A. debequaeus, A. desereticus, A. desperatus, A. deterior, A. detritalis, A. diaphanus, A. didymocarpus, A. distortus, A. diversifolius, A. douglasii, A. drabelliformis, A. drummondii, A. duchesnensis, A. eastwoodiae, A. egglestonii, A. emoryanus, A. endopterus, A. ensiformis, A. episcopus, A. equisolensis, A. eremiticus, A. ertterae, A. eucosmus, A. eurekensis, A. eurylobus, A. falcatus, A. feensis, A. filipes, A. flavus, A. flexuosus, A. fucatus, A. funereus, A. gambelianus, A. geyeri, A. gibbsii, A. giganteus, A. gilensis, A. gilmanii, A. gilviflorus, A. glycyphyllos, A. gracilis, A. grayi, A. gypsodes, A. hallii, A. hamiltonii, A. harrisonii, A. heilii, A. henrimontanensis, A. holmgreniorum, A. hoodianus, A. hornii, A. howellii, A. humillimus, A. humistratus, A. hyalinus, A. hypoxylus, A. inflexus, A. insularis, A. inversus, A. inyoensis, A. iodanthus, A. iodopetalus, A. iselyi, A. jaegerianus, A. jejunus, A. johannis-howellii, A. kelseyae, A. kerrii, A. knightii, A. laccoliticus, A. lancearius, A. laxmannii, A. layneae, A. leibergii, A. lemmonii, A. lentiformis, A. lentiginosus, A. leptaleus, A. leptocarpus, A. leucolobus, A. limnocharis, A. lindheimeri, A. linifolius, A. loanus, A. lonchocarpus, A. lotiflorus, A. lutosus, A. lyallii, A. macrodon, A. magdalenae, A. malacoides, A. malacus, A. megacarpus, A. michauxii, A. microcymbus, A. microcystis, A. microlobium, A. micromerius, A. miguelensis, A. minthorniae, A. misellus, A. miser, A. missouriensis, A. moencoppensis, A. mohavensis, A. mollissimus, A. molybdenus, A. monoensis, A. montii, A. monumentalis, A. mulfordiae, A. multiflorus, A. musiniensis, A. naturitensis, A. neglectus, A. nelsonianus, A. neomexicanus, A. nevinii, A. newberryi, A. nidularius, A. nothoxys, A. nudisiliquus, A. nutans, A. nutriosensis, A. nuttallianus, A. nuttallii, A. nutzotinensis, A. nyensis, A. obcordatus, A. obscurus, A. oniciformis, A. oocalycis, A. oocarpus, A. oophorus, A. oreganus, A. osterhoutii, A. oxyphysus, A. pachypus, A. palmeri, A. panamintensis, A. pardalinus, A. parryi, A. pattersonii, A. pauperculus, A. paysonii, A. peckii, A. pectinatus, A. perianus, A. phoenix, A. pictiformis, A. pinonis, A. piscator, A. piutensis, A. plattensis, A. platytropis, A. polaris, A. pomonensis, A. porrectus, A. praelongus, A. preussii, A. proimanthus, A. proximus, A. pseudiodanthus, A. pterocarpus, A. pubentissimus, A. pulsiferae, A. puniceus, A. purshii, A. pycnostachyus, A. racemosus, A. rafaelensis, A. rattanii, A. ravenii, A. recurvus, A. reflexus, A. remotus, A. reventiformis, A. reventus, A. riparius, A. ripleyi, A. robbinsii, A. rusbyi, A. sabulonum, A. sabulosus, A. salmonis, A. saurinus, A. scaphoides, A. schmolliae, A. sclerocarpus, A. scopulorum, A. sepultipes, A. serenoi, A. sericoleucus, A. serpens, A. sesquiflorus, A. sheldonii, A. shevockii, A. shortianus, A. siliceus, A. simplicifolius, A. sinuatus, A. solitarius, A. sophoroides, A. soxmaniorum, A. spaldingii, A. sparsiflorus, A. spatulatus, A. speirocarpus, A. straturensis, A. striatiflorus, A. subcinereus, A. subvestitus, A. succumbens, A. tegetarioides, A. tener, A. tennesseensis, A. tephrodes, A. terminalis, A. tetrapterus, A. thurberi, A. tibetanus, A. tidestromii, A. tiehmii, A. titanophilus, A. toanus, A. toquimanus, A. tortipes, A. traskiae, A. tricarinatus, A. trichopodus, A. tridactylicus, A. troglodytus, A. tweedyi, A. tyghensis, A. umbellatus, A. umbraticus, A. uncialis, A. utahensis, A. vaccarum, A. vallaris, A. vexilliflexus, A. villosus, A. wardii, A. waterfallii, A. webberi, A. welshii, A. wetherillii, A. whitneyi, A. williamsii, A. wingatanus, A. wittmannii, A. woodruffii, A. wootonii, A. wrightii, A. xiphoides, A. yoderwilliamsii, A. zionis |
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Kentrophyta montana |
Barneby: Mem. New York Bot. Gard. 13: 364. (1964) |
A. Gray: Proc. Acad. Nat. Sci. Philadelphia 15: 60. (1863) |
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