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Hayden's milkvetch

silver-leafed milkvetch, two-grooved milkvetch

Habit Plants stout, erect or diffuse and spreading, herbage usually pubescent. Plants 15–75 cm, mostly strigulose.
Stems

often erect or ascending, sometimes decumbent, diffuse, or spreading, strigulose or glabrous.

Leaves

3–13.5 cm;

stipules connate-sheathing at proximal nodes, connate or distinct at distal nodes, 2.5–10(–12) mm, submembranous becoming scarious, pallid;

leaflets (7–)11–35, blades lanceolate-oblong to oblong, elliptic, or oblanceolate, 5–27(–32) mm, apex usually acute or obtuse, rarely emarginate, surfaces strigulose abaxially, glabrous or glabrate adaxially.

Leaflets

(13–)21–35, 5–27 mm.

Racemes

35–80-flowered;

axis (4–)5.5–25 cm in fruit;

bracts (2.5–)3–5 mm.

(15–)25–80-flowered;

axis 3–25 cm in fruit;

bracts (1–)2.5–7 mm;

bracteoles 0–2.

Peduncles

erect, ascending, or incurved, 2.5–13 cm.

Pedicels

1–3.5 mm.

Flowers

8–11 mm;

calyx usually pallid, tube 3.1–4 mm, lobes 1–2.7 mm;

corolla white or whitish to ochroleucous;

banner longer than keel.

(6.7–)8–17.5 mm;

calyx obliquely campanulate, gibbous-saccate, 3.5–9.6 mm, sparsely strigose, tube 2.8–5.7(–7) mm, lobes subulate, (0.7–)1–4.5(–6) mm;

corolla ochroleucous, white, whitish, or pink-purple, keel tip often maculate, sometimes purple;

banner recurved through 45° (except 90° in var. nevadensis);

keel 7–13 mm.

Legumes

ellipsoid or oblong-ellipsoid, (5–)6.5–9.5 × 2–4 mm, transversely rugose-reticulate, strigulose;

stipe 1.4–3 mm.

stramineous, straight or slightly curved, linear, ellipsoid, or oblong-ellipsoid, dorsiventrally compressed, abaxial face with suture forming prominent ridge flanked by depressed grooves, adaxial face convexly rounded, (5–)6.5–17(–20) × 2–4.5 mm, fleshy becoming thinly leathery or papery, strigulose or glabrous;

stipe 1.4–5.2(–6) mm.

Seeds

5–8.

5–15.

2n

= 24.

Astragalus bisulcatus var. haydenianus

Astragalus bisulcatus

Phenology Flowering May–Jul.
Habitat Sagebrush-mountain brush, pinyon-juniper, ponderosa pine, and spruce-fir com­munities, on fine-textured, often saline, seleniferous substrates.
Elevation 1900–3300 m. (6200–10800 ft.)
Distribution
from FNA
AZ; CO; NM; UT; WY
[BONAP county map]
from USDA
w North America
[WildflowerSearch map]
[BONAP county map]
Discussion

Variety haydenianus from the Wasatch Plateau in Utah has been confused by some workers with the similarly pale-flowered var. major, but the flowers are consistently smaller, much more numerous, and the fruits smaller than in var. major.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 4 (4 in the flora).

Astragalus bisulcatus is an ill-scented primary indicator of selenium in most areas. In the Missouri drainage, it is easily recognized by its two-grooved fruits, and is relatively nonvariable and presents no taxonomic problems. West of the Front Ranges it is multivariate, the many forms difficult to classify.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Flowers 6.7–9.2 mm; corollas ochroleucous, ban­ner shorter than keel; raceme bracts 1–2.5 mm; ec Nevada.
var. nevadensis
1. Flowers 8–17.5 mm; corollas white, whitish, ochroleucous, or pink-purple, banner longer than keel; raceme bracts (2–)2.5–7 mm; widespread, not in Nevada.
→ 2
2. Legumes transversely rugose-reticulate, (5–)6.5–9.5 mm; flowers 8–11 mm; nw New Mexico, adjoining Arizona and Utah, north­ward through w Colorado to ne Utah and sw Wyoming.
var. haydenianus
2. Legumes smooth or faintly reticulate, 8–17(–20) mm; flowers 11–17.5 mm; Alberta to Manitoba, southward and westward to Arizona and Oklahoma.
→ 3
3. Flowers 13–17.5 mm; corollas usually pink-purple, rarely white; calyces usually red-purple; Alberta to Manitoba, southward and westward to Utah, New Mexico, and Oklahoma.
var. bisulcatus
3. Flowers 11–15 mm; corollas white or ochro­leucous, less commonly with purple center and purple lines, or suffused with pale purple throughout; calyces usually white; nw Colorado, sw Wyoming south­ward to c Utah and to n Arizona, disjunct in Fremont County, Idaho.
var. major
Source FNA vol. 11. FNA vol. 11.
Parent taxa Fabaceae > subfam. Faboideae > Astragalus > sect. Bisulcati > Astragalus bisulcatus Fabaceae > subfam. Faboideae > Astragalus > sect. Bisulcati
Sibling taxa
A. bisulcatus var. bisulcatus, A. bisulcatus var. major, A. bisulcatus var. nevadensis
A. accidens, A. accumbens, A. ackermanii, A. acutirostris, A. adanus, A. aequalis, A. agnicidus, A. agrestis, A. albens, A. albulus, A. allochrous, A. alpinus, A. altus, A. alvordensis, A. amblytropis, A. americanus, A. amnis-amissi, A. amphioxys, A. ampullarioides, A. ampullarius, A. andersonii, A. anisus, A. anserinus, A. applegatei, A. aquilonius, A. aretioides, A. argophyllus, A. aridus, A. arizonicus, A. arrectus, A. arthurii, A. asclepiadoides, A. asotinensis, A. asymmetricus, A. atratus, A. atropubescens, A. atwoodii, A. austiniae, A. australis, A. barnebyi, A. barrii, A. beathii, A. beatleyae, A. beckwithii, A. bernardinus, A. bibullatus, A. bicristatus, A. bodinii, A. bolanderi, A. bourgovii, A. brandegeei, A. brauntonii, A. brazoensis, A. breweri, A. californicus, A. callithrix, A. calycosus, A. camptopus, A. canadensis, A. caricinus, A. casei, A. castaneiformis, A. castetteri, A. ceramicus, A. cerussatus, A. chamaeleuce, A. chamaemeniscus, A. chinensis, A. chloödes, A. chuskanus, A. cibarius, A. cicer, A. cimae, A. claranus, A. clevelandii, A. cliffordii, A. cobrensis, A. coccineus, A. collinus, A. coltonii, A. columbianus, A. concordius, A. congdonii, A. conjunctus, A. consobrinus, A. contortuplicatus, A. convallarius, A. cottamii, A. crassicarpus, A. cremnophylax, A. cronquistii, A. crotalariae, A. curtipes, A. curvicarpus, A. cusickii, A. cutleri, A. cyaneus, A. cymboides, A. deanei, A. debequaeus, A. desereticus, A. desperatus, A. deterior, A. detritalis, A. diaphanus, A. didymocarpus, A. distortus, A. diversifolius, A. douglasii, A. drabelliformis, A. drummondii, A. duchesnensis, A. eastwoodiae, A. egglestonii, A. emoryanus, A. endopterus, A. ensiformis, A. episcopus, A. equisolensis, A. eremiticus, A. ertterae, A. eucosmus, A. eurekensis, A. eurylobus, A. falcatus, A. feensis, A. filipes, A. flavus, A. flexuosus, A. fucatus, A. funereus, A. gambelianus, A. geyeri, A. gibbsii, A. giganteus, A. gilensis, A. gilmanii, A. gilviflorus, A. glycyphyllos, A. gracilis, A. grayi, A. gypsodes, A. hallii, A. hamiltonii, A. harrisonii, A. heilii, A. henrimontanensis, A. holmgreniorum, A. hoodianus, A. hornii, A. howellii, A. humillimus, A. humistratus, A. hyalinus, A. hypoxylus, A. inflexus, A. insularis, A. inversus, A. inyoensis, A. iodanthus, A. iodopetalus, A. iselyi, A. jaegerianus, A. jejunus, A. johannis-howellii, A. kelseyae, A. kentrophyta, A. kerrii, A. knightii, A. laccoliticus, A. lancearius, A. laxmannii, A. layneae, A. leibergii, A. lemmonii, A. lentiformis, A. lentiginosus, A. leptaleus, A. leptocarpus, A. leucolobus, A. limnocharis, A. lindheimeri, A. linifolius, A. loanus, A. lonchocarpus, A. lotiflorus, A. lutosus, A. lyallii, A. macrodon, A. magdalenae, A. malacoides, A. malacus, A. megacarpus, A. michauxii, A. microcymbus, A. microcystis, A. microlobium, A. micromerius, A. miguelensis, A. minthorniae, A. misellus, A. miser, A. missouriensis, A. moencoppensis, A. mohavensis, A. mollissimus, A. molybdenus, A. monoensis, A. montii, A. monumentalis, A. mulfordiae, A. multiflorus, A. musiniensis, A. naturitensis, A. neglectus, A. nelsonianus, A. neomexicanus, A. nevinii, A. newberryi, A. nidularius, A. nothoxys, A. nudisiliquus, A. nutans, A. nutriosensis, A. nuttallianus, A. nuttallii, A. nutzotinensis, A. nyensis, A. obcordatus, A. obscurus, A. oniciformis, A. oocalycis, A. oocarpus, A. oophorus, A. oreganus, A. osterhoutii, A. oxyphysus, A. pachypus, A. palmeri, A. panamintensis, A. pardalinus, A. parryi, A. pattersonii, A. pauperculus, A. paysonii, A. peckii, A. pectinatus, A. perianus, A. phoenix, A. pictiformis, A. pinonis, A. piscator, A. piutensis, A. plattensis, A. platytropis, A. polaris, A. pomonensis, A. porrectus, A. praelongus, A. preussii, A. proimanthus, A. proximus, A. pseudiodanthus, A. pterocarpus, A. pubentissimus, A. pulsiferae, A. puniceus, A. purshii, A. pycnostachyus, A. racemosus, A. rafaelensis, A. rattanii, A. ravenii, A. recurvus, A. reflexus, A. remotus, A. reventiformis, A. reventus, A. riparius, A. ripleyi, A. robbinsii, A. rusbyi, A. sabulonum, A. sabulosus, A. salmonis, A. saurinus, A. scaphoides, A. schmolliae, A. sclerocarpus, A. scopulorum, A. sepultipes, A. serenoi, A. sericoleucus, A. serpens, A. sesquiflorus, A. sheldonii, A. shevockii, A. shortianus, A. siliceus, A. simplicifolius, A. sinuatus, A. solitarius, A. sophoroides, A. soxmaniorum, A. spaldingii, A. sparsiflorus, A. spatulatus, A. speirocarpus, A. straturensis, A. striatiflorus, A. subcinereus, A. subvestitus, A. succumbens, A. tegetarioides, A. tener, A. tennesseensis, A. tephrodes, A. terminalis, A. tetrapterus, A. thurberi, A. tibetanus, A. tidestromii, A. tiehmii, A. titanophilus, A. toanus, A. toquimanus, A. tortipes, A. traskiae, A. tricarinatus, A. trichopodus, A. tridactylicus, A. troglodytus, A. tweedyi, A. tyghensis, A. umbellatus, A. umbraticus, A. uncialis, A. utahensis, A. vaccarum, A. vallaris, A. vexilliflexus, A. villosus, A. wardii, A. waterfallii, A. webberi, A. welshii, A. wetherillii, A. whitneyi, A. williamsii, A. wingatanus, A. wittmannii, A. woodruffii, A. wootonii, A. wrightii, A. xiphoides, A. yoderwilliamsii, A. zionis
Subordinate taxa
A. bisulcatus var. bisulcatus, A. bisulcatus var. haydenianus, A. bisulcatus var. major, A. bisulcatus var. nevadensis
Synonyms A. haydenianus, A. bisulcatus subsp. haydenianus, A. grallator, Diholcos haydenianus, Tragacantha haydeniana Phaca bisulcata
Name authority (A. Gray) Barneby: Mem. New York Bot. Gard. 13: 413. (1964) (Hooker) A. Gray in War Department [U.S.]: Pacif. Railr. Rep. 12: 42. (1860)
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