Asteraceae (tribe Heliantheae) subtribe Helianthinae(synonym of Asteraceae tribe Heliantheae subtribe Helianthinae) |
Simsia |
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bush sunflower |
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Habit | Annuals, perennials, subshrubs, or shrubs, (2–)20–500 cm. | Annuals, perennials, or subshrubs [shrubs], 20–400 cm. | ||||||||||||||||||||||||||||||||||||||||
Stems | erect or ascending [decumbent], sparingly to freely branched. |
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Leaves | usually mostly cauline, rarely mostly basal; mostly opposite (distal often alternate) or mostly alternate, rarely whorled; petiolate or sessile; blades (often 3- or 5-nerved) orbiculate or deltate to lanceolate or linear (and intermediate shapes), sometimes lobed, ultimate margins entire or toothed, faces usually hairy (often hispid to scabrellous), often gland-dotted. |
cauline; opposite (proximal) or alternate [whorled]; petiolate (petioles often ± winged, often with expanded bases, those bases sometimes fused to form nodal “discs”) [sessile]; blades 3-nerved from bases, mostly deltate to ovate [linear], sometimes 3- [5-]lobed[pinnatifid], bases cordate to cuneate, ultimate margins entire or toothed, faces hirsute, hispid, pilose, puberulent, scabrous, or scabro-hispid [sericeous], often gland-dotted or ± stipitate-glandular to glandular-puberulent. |
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Involucres | cylindric to hemispheric or rotate. |
campanulate [ovoid-campanulate to urceolate], 5–16[–22] mm diam. |
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Receptacles | usually flat or convex, sometimes hemispheric or conic, usually paleate (epaleate in Lagascea; paleae usually falling, sometimes persistent, mostly oblong to linear, often conduplicate, herbaceous to scarious, apices sometimes ± 3-lobed or -toothed; becoming hardened perigynia completely investing cypselae in Sclerocarpus). |
low-convex, paleate (paleae conduplicate, ± enclosing cypselae). |
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Ray florets | 0 or 5–30(–100+), neuter, or styliferous and sterile; corollas usually yellow to orange, sometimes partly or wholly brown to purple or red-brown to reddish. |
[0–]5–21[–45], styliferous and sterile; corollas orange-yellow [lemon-yellow, pink, purple, or white]. |
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Disc florets | 1 (Lagascea) or 10–200(–1000+), bisexual, fertile; corollas usually yellow to orange, sometimes brown to purple or reddish to red-brown, tubes shorter than or about equaling cylindric or funnelform throats, lobes 5, ± deltate (equal); anther thecae dark or pale; stigmatic papillae usually continuous. |
[12–]13–154[–172], bisexual, fertile; corollas concolorous with rays (usually turning purple apically), tubes (often glandular-hairy) shorter than throats, lobes 5, ± triangular (anthers black, yellow, or yellow proximally and bronze or purple distally; style branches relatively slender, apices sometimes attenuate). |
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Phyllaries | persistent, 4–45(–100+) in 1–6+ series (usually distinct, ± connate in Lagascea, broadly ovate to linear, subequal or unequal). |
persistent, [11–]13–43[–66] in 2–4 series (tightly appressed to broadly reflexed, unequal to subequal). |
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Calyculi | 0. |
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Heads | usually radiate, sometimes discoid, borne singly or in corymbiform (paniculiform or racemiform) arrays (glomerules or second-order heads of 1–2-flowered heads in Lagascea). |
radiate [discoid], borne singly or in 2s or 3s, or in tight to loose, corymbiform [paniculiform] arrays. |
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Cypselae | often ± compressed, flattened, biconvex, clavate, columnar, obovoid, obpyramidal, or prismatic (lengths usually 1–2 times diams.), glabrous or hairy (faces and/or angles); pappi 0, or (often readily falling or fragile, sometimes persistent) usually of (1–)2(–8+) (sometimes aristate) scales and/or awns, sometimes coroniform. |
flattened, thin-margined [thickened, biconvex] (shoulders minute to conspicuous, faces glabrous or hairy); pappi 0, or fragile or readily falling, of 2 ± subulate scales [plus 4–12 shorter scales]. |
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x | = 17. |
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Asteraceae (tribe Heliantheae) subtribe Helianthinae |
Simsia |
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Distribution | Mostly subtropical; tropical; and warm-temperate New World |
sw United States; Mexico; Central America; South America; West Indies (Jamaica) |
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Discussion | Genera 17, species 363 (8 genera, 69 species, including 1 hybrid, in the flora). Circumscription of Helianthinae adopted here (i.e., that of H. Robinson 1981) is narrower than traditional circumscriptions (e.g., T. F. Stuessy 1977[1978]), which included genera here treated in Ecliptinae, Galinsoginae, and Rudbeckiinae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 20 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 135. | FNA vol. 21, p. 140. | ||||||||||||||||||||||||||||||||||||||||
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Synonyms | subtribe Lagasceinae | |||||||||||||||||||||||||||||||||||||||||
Name authority | Cassini ex Dumortier: Fl. Belg., 71. (1827) | Persoon: Syn. Pl. 2: 478. (1807) | ||||||||||||||||||||||||||||||||||||||||
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