Asteraceae (tribe Heliantheae) subtribe Helianthinae(synonym of Asteraceae tribe Heliantheae subtribe Helianthinae) |
Sclerocarpus |
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bonebract, Mexican bone-bract |
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Habit | Annuals, perennials, subshrubs, or shrubs, (2–)20–500 cm. | Annuals [perennials or subshrubs], 60–200 cm. | ||||||||||||||||||||||||||||||||||||
Stems | sprawling to erect, branched from bases or ± throughout. |
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Leaves | usually mostly cauline, rarely mostly basal; mostly opposite (distal often alternate) or mostly alternate, rarely whorled; petiolate or sessile; blades (often 3- or 5-nerved) orbiculate or deltate to lanceolate or linear (and intermediate shapes), sometimes lobed, ultimate margins entire or toothed, faces usually hairy (often hispid to scabrellous), often gland-dotted. |
mostly cauline; opposite (proximal) or alternate; petiolate [sessile]; blades 3-nerved, ovate to rhombic or lanceolate, bases cuneate to rounded, margins coarsely toothed, faces glabrous or ± scabrous. |
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Involucres | cylindric to hemispheric or rotate. |
rotate to hemispheric, 12–20 mm diam. |
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Receptacles | usually flat or convex, sometimes hemispheric or conic, usually paleate (epaleate in Lagascea; paleae usually falling, sometimes persistent, mostly oblong to linear, often conduplicate, herbaceous to scarious, apices sometimes ± 3-lobed or -toothed; becoming hardened perigynia completely investing cypselae in Sclerocarpus). |
low-conic, paleate (paleae each closely investing subtended floret, each forming a hard perigynium around a fruit and shed with it). |
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Ray florets | 0 or 5–30(–100+), neuter, or styliferous and sterile; corollas usually yellow to orange, sometimes partly or wholly brown to purple or red-brown to reddish. |
5–8, neuter; corollas (readily falling) yellow to orange. |
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Disc florets | 1 (Lagascea) or 10–200(–1000+), bisexual, fertile; corollas usually yellow to orange, sometimes brown to purple or reddish to red-brown, tubes shorter than or about equaling cylindric or funnelform throats, lobes 5, ± deltate (equal); anther thecae dark or pale; stigmatic papillae usually continuous. |
10–50, bisexual, fertile; corollas usually yellow to orange, sometimes with purple, tubes much shorter than cylindric throats, lobes 5, lance-linear. |
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Phyllaries | persistent, 4–45(–100+) in 1–6+ series (usually distinct, ± connate in Lagascea, broadly ovate to linear, subequal or unequal). |
persistent, 5–8[–21] in 1(–2) series (green, oblong to linear). |
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Calyculi | 0. |
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Heads | usually radiate, sometimes discoid, borne singly or in corymbiform (paniculiform or racemiform) arrays (glomerules or second-order heads of 1–2-flowered heads in Lagascea). |
radiate, borne singly. |
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Cypselae | often ± compressed, flattened, biconvex, clavate, columnar, obovoid, obpyramidal, or prismatic (lengths usually 1–2 times diams.), glabrous or hairy (faces and/or angles); pappi 0, or (often readily falling or fragile, sometimes persistent) usually of (1–)2(–8+) (sometimes aristate) scales and/or awns, sometimes coroniform. |
(within perigynia) weakly compressed, obliquely obovoid or arcuate (glabrous); pappi 0, or coroniform (of distinct or connate scales or bristles). |
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x | = 12. |
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Asteraceae (tribe Heliantheae) subtribe Helianthinae |
Sclerocarpus |
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Distribution | Mostly subtropical; tropical; and warm-temperate New World |
Mexico; Central America; s United States; one species in Old World tropics |
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Discussion | Genera 17, species 363 (8 genera, 69 species, including 1 hybrid, in the flora). Circumscription of Helianthinae adopted here (i.e., that of H. Robinson 1981) is narrower than traditional circumscriptions (e.g., T. F. Stuessy 1977[1978]), which included genera here treated in Ecliptinae, Galinsoginae, and Rudbeckiinae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 12 (1 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 135. | FNA vol. 21, p. 137. | ||||||||||||||||||||||||||||||||||||
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Synonyms | subtribe Lagasceinae | |||||||||||||||||||||||||||||||||||||
Name authority | Cassini ex Dumortier: Fl. Belg., 71. (1827) | Jacquin: Icon. Pl. Rar. 1: 17, plate 176. (1781) | ||||||||||||||||||||||||||||||||||||
Web links |