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acuate, doll's-head, lagascea, silk-leaf, velvet-bush

Habit Annuals, perennials, subshrubs, or shrubs, (2–)20–500 cm. Shrubs [annuals, perennials], to 50–100[–300] cm.
Stems

erect [ascending to decumbent], branched from bases or ± throughout.

Leaves

usually mostly cauline, rarely mostly basal; mostly opposite (distal often alternate) or mostly alternate, rarely whorled;

petiolate or sessile;

blades (often 3- or 5-nerved) orbiculate or deltate to lanceolate or linear (and intermediate shapes), sometimes lobed, ultimate margins entire or toothed, faces usually hairy (often hispid to scabrellous), often gland-dotted.

cauline; opposite;

petiolate [sessile];

blades 3-nerved, lance-ovate to ovate [lanceolate to oblanceolate], bases broadly cuneate [to subauriculate], margins ± serrate, faces usually sericeous to strigose or glabrate, often stipitate-glandular.

Involucres

cylindric to hemispheric or rotate.

cylindric, 1–2 mm diam. (glomerules of heads usually subtended by leaves or ± foliaceous bracts).

Receptacles

usually flat or convex, sometimes hemispheric or conic, usually paleate (epaleate in Lagascea; paleae usually falling, sometimes persistent, mostly oblong to linear, often conduplicate, herbaceous to scarious, apices sometimes ± 3-lobed or -toothed; becoming hardened perigynia completely investing cypselae in Sclerocarpus).

convex (often hirtellous), rarely paleate (paleae linear).

Ray florets

0 or 5–30(–100+), neuter, or styliferous and sterile;

corollas usually yellow to orange, sometimes partly or wholly brown to purple or red-brown to reddish.

0.

Disc florets

1 (Lagascea) or 10–200(–1000+), bisexual, fertile;

corollas usually yellow to orange, sometimes brown to purple or reddish to red-brown, tubes shorter than or about equaling cylindric or funnelform throats, lobes 5, ± deltate (equal);

anther thecae dark or pale;

stigmatic papillae usually continuous.

1(–2)[–8], bisexual, fertile;

corollas yellow [white, pink, or red], tubes shorter than cylindric to campanulate throats, lobes 5, lance-linear to lance-ovate [rounded-deltate] (often hairy).

Phyllaries

persistent, 4–45(–100+) in 1–6+ series (usually distinct, ± connate in Lagascea, broadly ovate to linear, subequal or unequal).

persistent, 4–5[–8+] in ± 1 series (linear-attenuate, proximally connate, often 1 or more with 1[–3+] glands in abaxial face).

Calyculi

0.

Heads

usually radiate, sometimes discoid, borne singly or in corymbiform (paniculiform or racemiform) arrays (glomerules or second-order heads of 1–2-flowered heads in Lagascea).

discoid, borne in headlike glomerules (of [8–]30–50+, 1(–2)[–8]-flowered heads, glomerules borne singly or in ± corymbiform [racemiform] arrays).

Cypselae

often ± compressed, flattened, biconvex, clavate, columnar, obovoid, obpyramidal, or prismatic (lengths usually 1–2 times diams.), glabrous or hairy (faces and/or angles);

pappi 0, or (often readily falling or fragile, sometimes persistent) usually of (1–)2(–8+) (sometimes aristate) scales and/or awns, sometimes coroniform.

(brown to black) narrowly cylindric to obovoid or clavate (faces glabrous or pilosulous, minutely grooved);

pappi ± coroniform.

x

= 17.

Asteraceae (tribe Heliantheae) subtribe Helianthinae

Lagascea

Distribution
Mostly subtropical; tropical; and warm-temperate New World
from USDA
sw United States; Mexico; Central America (to Nicaragua) [Introduced in West Indies, South America, Asia (India, Java, Sri Lanka, Thailand), Africa, Pacific Islands (Hawaii)]
[BONAP county map]
Discussion

Genera 17, species 363 (8 genera, 69 species, including 1 hybrid, in the flora).

Circumscription of Helianthinae adopted here (i.e., that of H. Robinson 1981) is narrower than traditional circumscriptions (e.g., T. F. Stuessy 1977[1978]), which included genera here treated in Ecliptinae, Galinsoginae, and Rudbeckiinae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 8 (1 in the flora).

Lagascea mollis Cavanilles, now a nearly pantropical weed, was collected at Apalachicola, Franklin County, Florida (Chapman, n.d., presumably ca. 1860), evidently from an ephemeral population, probably from ballast.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Phyllaries 4–5 in ± 1 series (connate); disc florets 1(–2; the 1–2-flowered heads aggregated in glomerules or second-order heads)
Lagascea
1. Phyllaries 5–45(–100+) in (1–)2–6+ series (distinct); disc florets (10–)15–200(–1000+)
→ 2
2. Receptacles paleate (each palea completely investing and falling with a cypsela, eachforming a hardened perigynium)
Sclerocarpus
2. Receptacles paleate (paleae sometimes conduplicate, ± enfolding cypselae, not forming perigynia)
→ 3
3. Heads borne singly (peduncles usually distally dilated, fistulose)
Tithonia
3. Heads borne singly or in corymbiform, paniculiform, racemiform, or thyrsiform arrays (peduncles rarely, if ever, notably dilated or fistulose)
→ 4
4. Cypselae flattened, thin-margined
Simsia
4. Cypselae ± compressed, biconvex, or 3- or 4-angled, often obpyramidal
→ 5
5. Cypselae glabrous or glabrate; pappi 0
→ 6
5. Cypselae usually ± strigose, sometimes glabrous or glabrate; pappi of 2–6+, persistent, readily falling, or tardily falling scales
→ 8
6. Shrubs (leaves often lobed, lobes usually 3–9, ± linear)
Viguiera
6. Annuals or perennials (leaves not lobed)
→ 7
7. Annuals; leaf blades lanceolate to linear; involucres 5–6 mm diam.;phyllaries 11–17
Helianthus
7. Annuals or perennials; leaf blades lance-linear, lanceolate, ovate, rhombic, or rhombic-ovate; involucres 6–14 mm diam.; phyllaries14–25
Heliomeris
8. Shrubs; involucres 5–9 mm diam
Bahiopsis
8. Annuals or perennials; involucres (5–)7–40+ mm diam
→ 9
9. Pappi readily falling
Helianthus
9. Pappi persistent or tardily falling
Viguiera
Source FNA vol. 21, p. 135. FNA vol. 21, p. 136. Treatment author: Elizabeth M. Harris.
Parent taxa Asteraceae > tribe Heliantheae Asteraceae > tribe Heliantheae > subtribe Helianthinae
Subordinate taxa
Bahiopsis, Helianthus, Heliomeris, Lagascea, Sclerocarpus, Simsia, Tithonia, Viguiera
L. decipiens
Synonyms subtribe Lagasceinae
Name authority Cassini ex Dumortier: Fl. Belg., 71. (1827) Cavanilles: Anales Ci. Nat. 6: 331. (1803)
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