Asteraceae (tribe Heliantheae) subtribe Helianthinae(synonym of Asteraceae tribe Heliantheae subtribe Helianthinae) |
Heliomeris |
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false goldeneye, golden-eye |
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Habit | Annuals, perennials, subshrubs, or shrubs, (2–)20–500 cm. | Annuals or perennials, (2–)10–90(–120+) cm. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Stems | erect, branched distally or ± throughout. |
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Leaves | usually mostly cauline, rarely mostly basal; mostly opposite (distal often alternate) or mostly alternate, rarely whorled; petiolate or sessile; blades (often 3- or 5-nerved) orbiculate or deltate to lanceolate or linear (and intermediate shapes), sometimes lobed, ultimate margins entire or toothed, faces usually hairy (often hispid to scabrellous), often gland-dotted. |
mostly cauline; opposite or alternate; sessile or subsessile; blades usually 1(–3)-nerved, elliptic, lance-linear, lanceolate, lance-ovate, linear, ovate, rhombic, or rhombic-ovate, margins entire (often revolute), faces hispid or strigose to stigillose, sometimes gland-dotted. |
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Involucres | cylindric to hemispheric or rotate. |
hemispheric or broader, 6–14 mm diam. |
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Receptacles | usually flat or convex, sometimes hemispheric or conic, usually paleate (epaleate in Lagascea; paleae usually falling, sometimes persistent, mostly oblong to linear, often conduplicate, herbaceous to scarious, apices sometimes ± 3-lobed or -toothed; becoming hardened perigynia completely investing cypselae in Sclerocarpus). |
conic; paleate (paleae tan to brown, ovate to oblong-rectangular, conduplicate). |
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Ray florets | 0 or 5–30(–100+), neuter, or styliferous and sterile; corollas usually yellow to orange, sometimes partly or wholly brown to purple or red-brown to reddish. |
5–15, neuter; corollas yellow (laminae elliptic, oblong, obovate, oval, or ovate). |
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Disc florets | 1 (Lagascea) or 10–200(–1000+), bisexual, fertile; corollas usually yellow to orange, sometimes brown to purple or reddish to red-brown, tubes shorter than or about equaling cylindric or funnelform throats, lobes 5, ± deltate (equal); anther thecae dark or pale; stigmatic papillae usually continuous. |
25–50+, bisexual, fertile; corollas yellow, tubes shorter than campanulate throats, lobes 5, triangular (style branches relatively slender, apices acute). |
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Phyllaries | persistent, 4–45(–100+) in 1–6+ series (usually distinct, ± connate in Lagascea, broadly ovate to linear, subequal or unequal). |
persistent, 14–25 in ± 2–3 series (mostly lance-linear, herbaceous). |
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Calyculi | 0. |
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Heads | usually radiate, sometimes discoid, borne singly or in corymbiform (paniculiform or racemiform) arrays (glomerules or second-order heads of 1–2-flowered heads in Lagascea). |
radiate, borne singly or in cymiform to paniculiform arrays. |
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Cypselae | often ± compressed, flattened, biconvex, clavate, columnar, obovoid, obpyramidal, or prismatic (lengths usually 1–2 times diams.), glabrous or hairy (faces and/or angles); pappi 0, or (often readily falling or fragile, sometimes persistent) usually of (1–)2(–8+) (sometimes aristate) scales and/or awns, sometimes coroniform. |
(black, mottled, or gray-striate) weakly 4-angled, ± obpyramidal (glabrous); pappi 0. |
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x | = 8. |
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Asteraceae (tribe Heliantheae) subtribe Helianthinae |
Heliomeris |
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Distribution | Mostly subtropical; tropical; and warm-temperate New World |
sw United States; Mexico |
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Discussion | Genera 17, species 363 (8 genera, 69 species, including 1 hybrid, in the flora). Circumscription of Helianthinae adopted here (i.e., that of H. Robinson 1981) is narrower than traditional circumscriptions (e.g., T. F. Stuessy 1977[1978]), which included genera here treated in Ecliptinae, Galinsoginae, and Rudbeckiinae. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 5 (4 in the flora). Heliomeris has often been submerged within Viguiera; herbaceous phyllaries, epappose cypselae, and distinctive chromosome base number provide morphologic and genetic features that correlate with molecular phylogenetic studies to suggest that it be recognized as distinct. A species from granite outcrops of the southeastern United States, Helianthus porteri, shows remarkable morphologic similarity to Heliomeris, but cytologic, crossing, and molecular phylogenetic studies have provided abundant evidence that it is properly placed in Helianthus. W. F. Yates and C. B. Heiser (1979) provided a useful summary of Heliomeris, and their treatment is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 21, p. 135. | FNA vol. 21, p. 169. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Parent taxa | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Synonyms | subtribe Lagasceinae | Viguiera section H. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Name authority | Cassini ex Dumortier: Fl. Belg., 71. (1827) | Nuttall: Proc. Acad. Nat. Sci. Philadelphia 4: 19. (1848) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
Web links |