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Habit Annuals, perennials, subshrubs, or shrubs, (2–)20–500 cm. Shrubs, 50–150 cm.
Stems

ascending to erect, much branched.

Leaves

usually mostly cauline, rarely mostly basal; mostly opposite (distal often alternate) or mostly alternate, rarely whorled;

petiolate or sessile;

blades (often 3- or 5-nerved) orbiculate or deltate to lanceolate or linear (and intermediate shapes), sometimes lobed, ultimate margins entire or toothed, faces usually hairy (often hispid to scabrellous), often gland-dotted.

mostly cauline; alternate or opposite;

petiolate or sessile;

blades usually 3-nerved (from at or near bases), deltate, deltate-ovate, lanceolate, lance-ovate, or ovate, margins entire or toothed to laciniate, faces hispid, sericeous, or strigillose, often resinous or gland-dotted.

Involucres

cylindric to hemispheric or rotate.

campulate to hemispheric, (9–14 ×) 4–7 mm.

Receptacles

usually flat or convex, sometimes hemispheric or conic, usually paleate (epaleate in Lagascea; paleae usually falling, sometimes persistent, mostly oblong to linear, often conduplicate, herbaceous to scarious, apices sometimes ± 3-lobed or -toothed; becoming hardened perigynia completely investing cypselae in Sclerocarpus).

convex, paleate (paleae tan, conduplicate, apices acute).

Ray florets

0 or 5–30(–100+), neuter, or styliferous and sterile;

corollas usually yellow to orange, sometimes partly or wholly brown to purple or red-brown to reddish.

8–15, neuter;

corollas yellow (2–3-lobed).

Disc florets

1 (Lagascea) or 10–200(–1000+), bisexual, fertile;

corollas usually yellow to orange, sometimes brown to purple or reddish to red-brown, tubes shorter than or about equaling cylindric or funnelform throats, lobes 5, ± deltate (equal);

anther thecae dark or pale;

stigmatic papillae usually continuous.

40–50+, bisexual, fertile;

corollas yellow, tubes shorter than campanulate throats, lobes 5, triangular (style branches relatively slender, apices acute).

Phyllaries

persistent, 4–45(–100+) in 1–6+ series (usually distinct, ± connate in Lagascea, broadly ovate to linear, subequal or unequal).

persistent, 16–28 in 2–3 series (unequal, bases ovate to lance-ovate, indurate, apices abruptly narrowed, herbaceous).

Calyculi

0.

Heads

usually radiate, sometimes discoid, borne singly or in corymbiform (paniculiform or racemiform) arrays (glomerules or second-order heads of 1–2-flowered heads in Lagascea).

radiate, borne singly or (3–25+) in ± thyrsiform arrays.

Cypselae

often ± compressed, flattened, biconvex, clavate, columnar, obovoid, obpyramidal, or prismatic (lengths usually 1–2 times diams.), glabrous or hairy (faces and/or angles);

pappi 0, or (often readily falling or fragile, sometimes persistent) usually of (1–)2(–8+) (sometimes aristate) scales and/or awns, sometimes coroniform.

(brown to black) ± compressed, ± 3- or 4- angled, often obpyramidal, ± strigose;

pappi persistent, of 2(–6) lacerate, aristate scales (1–2.8 mm) plus (0–)2–6 lacerate scales (0.2–1 mm).

x

= 18.

Asteraceae (tribe Heliantheae) subtribe Helianthinae

Bahiopsis

Distribution
Mostly subtropical; tropical; and warm-temperate New World
sw United States; nw Mexico
[BONAP county map]
Discussion

Genera 17, species 363 (8 genera, 69 species, including 1 hybrid, in the flora).

Circumscription of Helianthinae adopted here (i.e., that of H. Robinson 1981) is narrower than traditional circumscriptions (e.g., T. F. Stuessy 1977[1978]), which included genera here treated in Ecliptinae, Galinsoginae, and Rudbeckiinae.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 12 (3 in the flora).

Although traditionally included within Viguiera, the species of Bahiopsis differ by their distinctive base chromosome number and form a basally diverging clade in subtribe Helianthinae, based on molecular data. Morphologically, they are most similar to Calanticaria, another segregate of Viguiera that is Mexican in geographic distribution, from which they differ by their branched capitulescences and having phyllaries in which the apical herbaceous portions are longer.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Phyllaries 4–5 in ± 1 series (connate); disc florets 1(–2; the 1–2-flowered heads aggregated in glomerules or second-order heads)
Lagascea
1. Phyllaries 5–45(–100+) in (1–)2–6+ series (distinct); disc florets (10–)15–200(–1000+)
→ 2
2. Receptacles paleate (each palea completely investing and falling with a cypsela, eachforming a hardened perigynium)
Sclerocarpus
2. Receptacles paleate (paleae sometimes conduplicate, ± enfolding cypselae, not forming perigynia)
→ 3
3. Heads borne singly (peduncles usually distally dilated, fistulose)
Tithonia
3. Heads borne singly or in corymbiform, paniculiform, racemiform, or thyrsiform arrays (peduncles rarely, if ever, notably dilated or fistulose)
→ 4
4. Cypselae flattened, thin-margined
Simsia
4. Cypselae ± compressed, biconvex, or 3- or 4-angled, often obpyramidal
→ 5
5. Cypselae glabrous or glabrate; pappi 0
→ 6
5. Cypselae usually ± strigose, sometimes glabrous or glabrate; pappi of 2–6+, persistent, readily falling, or tardily falling scales
→ 8
6. Shrubs (leaves often lobed, lobes usually 3–9, ± linear)
Viguiera
6. Annuals or perennials (leaves not lobed)
→ 7
7. Annuals; leaf blades lanceolate to linear; involucres 5–6 mm diam.;phyllaries 11–17
Helianthus
7. Annuals or perennials; leaf blades lance-linear, lanceolate, ovate, rhombic, or rhombic-ovate; involucres 6–14 mm diam.; phyllaries14–25
Heliomeris
8. Shrubs; involucres 5–9 mm diam
Bahiopsis
8. Annuals or perennials; involucres (5–)7–40+ mm diam
→ 9
9. Pappi readily falling
Helianthus
9. Pappi persistent or tardily falling
Viguiera
1. Leaves lanceolate to lance-ovate, margins ± laciniate, abaxial faces strigillose (resinous, shiny)
B. laciniata
1. Leaves deltate, deltate-ovate, or ovate, margins entire or serrate (not incised or laciniate-toothed), abaxial faces hispid, or sericeous (not shiny)
→ 2
2. Leaf blades 1–3.5 cm, margins usually toothed, faces: abaxial hispid and gland-dotted (and slightly reticulate), adaxial scabrous (bases of hairs notably enlarged)
B. parishii
2. Leaf blades 2.8–9 cm, margins entire, faces: abaxial sericeous and gland-dotted (andstrongly reticulate, not shiny), adaxial sericeous (hair bases slightly enlarged)
B. reticulata
Source FNA vol. 21, p. 135. FNA vol. 21, p. 174. Treatment author: Edward E. Schilling.
Parent taxa Asteraceae > tribe Heliantheae Asteraceae > tribe Heliantheae > subtribe Helianthinae
Subordinate taxa
Bahiopsis, Helianthus, Heliomeris, Lagascea, Sclerocarpus, Simsia, Tithonia, Viguiera
B. laciniata, B. parishii, B. reticulata
Synonyms subtribe Lagasceinae
Name authority Cassini ex Dumortier: Fl. Belg., 71. (1827) Kellogg: Proc. Calif. Acad. Sci. 2: 35. (1863)
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