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gallant soldier, garden pest, quickweed

Habit Annuals, perennials, subshrubs, or shrubs, (2–)10–150[–300+] cm. Annuals, 2–62 cm.
Stems

erect.

Leaves

mostly basal or mostly cauline; usually opposite (distal sometimes alternate);

petiolate or sessile;

blades (often 3- or 5-nerved) mostly deltate, elliptic, lanceolate, lance-ovate, linear, ovate, or rhombic, sometimes pinnately or palmately lobed, ultimate margins entire or ± toothed, faces glabrate, hispid, hispidulous, pilose, puberulent, scabrellous, scabrous, strigillose, or strigose, often gland-dotted or stipitate-glandular.

cauline; opposite;

petiolate;

blades (3-nerved) lanceolate to broadly ovate, margins entire or serrulate to serrate, faces glabrate to densely pilose.

Involucres

campanulate, cylindric, hemispheric, or ± obpyramidal.

hemispheric to campanulate, 2.5–6 mm diam.

Receptacles

flat, convex, or conic, paleate (paleae persistent or falling with cypselae, mostly oblong to lanceolate or linear, flat to conduplicate, herbaceous or scarious, often 2- or 3-lobed or -toothed).

conic, paleate (paleae persistent or falling, scarious, proximal broadly elliptic to obovate, often connate at bases or nearly to apices, united in groups of 2–3 to adjacent proximal phyllary, each complex often enclosing and shed with a ray cypsela; distal persistent or falling, lanceolate to obovate, entire or 2- or 3-lobed, convex to conduplicate).

Ray florets

0, or (3–)6–21, pistillate, fertile;

corollas yellow or whitish or pinkish [purplish].

[0–](3–)5(–8)[–15], pistillate, fertile;

corollas white or dull white to pinkish [purplish] (tubes pilose, laminae quadrate-obovate to oblong, lobes 0–3).

Disc florets

5–150+, bisexual, fertile (peripheral sometimes pistillate in Bebbia);

corollas yellow to orange, or whitish, pinkish, or purplish, tubes shorter than throats, lobes 5, deltate to lance-deltate or lanceolate (± equal [outer larger, e.g., some plants of Tridax]);

anther thecae pale or slightly darkened;

stigmatic papillae in 2 lines.

5–50[–150], bisexual, fertile;

corollas yellow, tubes (pilose) shorter than cylindric throats, lobes 5, deltate (anthers yellow; style-branch apices acute).

Phyllaries

persistent or falling, 6–30+ in 2–5+ series, usually distinct, elliptic, lanceolate, lance-linear, lance-ovate, oblong, or ovate, subequal or unequal (outer longer or shorter).

persistent or falling, 6–9[–16] in 2[–3] series (elliptic, lance-ovate, oblong, or ovate, outer shorter, herbaceous or scarious, margins entire or minutely laciniate).

Calyculi

0.

Heads

radiate or discoid (sometimes ± disciform in Bebbia), borne singly or in loose to tight, corymbiform or cymiform arrays.

radiate or discoid, in ± cymiform arrays.

Cypselae

± compressed, often clavate, obconic, or ovoid, or obpyramidal and 3-, 4-, or 5-angled, glabrous or sparsely to densely piloso-sericeous, sericeous, strigillose, or strigose;

pappi 0, or persistent, of 1–10+ subulate to acerose scales, or 5–20 ± spatulate, entire to erose, fimbriate, or laciniate, sometimes aristate, scales in 1 series, or 15–25+[–40], plumose, setiform scales (or flattened bristles) in 1 series.

obconic to obpyramidal, glabrous or strigose (rays often shed with subtending phyllary plus 2–3 adjacent paleae);

pappi 0, or persistent, of 5–20, white or gray, fimbriate, sometimes aristate scales.

x

= 8.

Asteraceae (tribe Heliantheae) subtribe Galinsoginae

Galinsoga

Distribution
Subtropical; tropical; and warm-temperate New World
from USDA
North America; Mexico; Central America; South America; West Indies; Bermuda [Introduced in Europe, Asia, Africa, Atlantic Islands, Pacific Islands, Australia]
[BONAP county map]
Discussion

Genera 15, species 103 (4 genera, 8 species in the flora).

All genera of Galinsoginae are centered in tropical and subtropical to warm-temperate North America and South America; relatively few extend into cool-temperate areas; fewer still are adventive in the Old World.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 15–33+ (2 in the flora).

Galinsoga is closely related to Sabazia Cassini of Mexico and South America and to at least some members of Alloispermum Willdenow, primarily from South America. Galinsoga might best be treated as a single larger genus including Alloispermum and Sabazia (J. M. Canne 1978; J. L. Panero et al. 1999).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Phyllaries 10–20 in ± 2 series (outer 4 broadly lanceolate, foliaceous, notably larger than inner)
Tetragonotheca
1. Phyllaries 6–30+ in 2–5 series (subequal or unequal, outer smaller than inner).
→ 2
2. Annuals (ray cypselae often each shed together with subtending phyllary and 2 adjacent paleae)
Galinsoga
2. Perennials, subshrubs, or shrubs (cypselae shed separate from paleae)
→ 3
3. Subshrubs or shrubs, 50–140 cm (often forming dense, rounded masses)
Bebbia
3. Perennials, 10–40 cm (stems procumbent to ascending)
Tridax
1. Phyllaries persistent; inner paleae 3-lobed, lobes 1/3+ total lengths, acute; disc pappi 0, or of 15–20 often gray, sometimes white, linear, fimbriate, obtuse scales 0.5–2 mm
G. parviflora
1. Phyllaries falling; inner paleae entire or 2- or 3-lobed, lobes to 1/3 total lengths, blunt; disc pappi 0, or of 1–5 or 14–20 white, lanceolate to oblanceolate, fimbriate, sometimes aristate, scales
G. quadriradiata
Source FNA vol. 21, p. 176. FNA vol. 21, p. 180. Treatment author: Judith M. Canne-Hilliker.
Parent taxa Asteraceae > tribe Heliantheae Asteraceae > tribe Heliantheae > subtribe Galinsoginae
Subordinate taxa
Bebbia, Galinsoga, Tetragonotheca, Tridax
G. parviflora, G. quadriradiata
Name authority Bentham & Hooker f.: Gen. Pl. 2: 198. (1873) Ruiz & Pavón: Fl. Peruv. Prodr., 110, plate 24. (1794)
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