Asteraceae (tribe Heliantheae) subtribe Coreopsidinae(synonym of Asteraceae tribe Heliantheae subtribe Coreopsidinae) |
Dicranocarpus |
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dicranocarpus, pitchfork |
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| Habit | Annuals, perennials, subshrubs, or shrubs, 10–400 cm (sometimes rhizomatous or with cormiform bases, stoloniferous in Coreopsis auriculata). | Annuals, 10–70 cm (± succulent; taprooted). | ||||||||||||||||||||||||||||
| Stems | 1, erect, branched dichotomously, divaricate (drying brittle). |
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| Leaves | usually cauline (sometimes mostly basal); usually mostly opposite (distal sometimes alternate), rarely whorled; petiolate or sessile; blades (often pinnately or palmately lobed, sometimes compound), or lobes or leaflets, mostly deltate, elliptic, filiform, lanceolate, linear, oblanceolate, or ovate, ultimate margins entire or toothed, faces usually glabrous or glabrate, sometimes hairy (rarely, if ever, gland-dotted). |
mostly cauline; opposite; obscurely petiolate; blades 1(–2)-pedately or -pinnately lobed (lobes filiform), ultimate margins entire, faces glabrous. |
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| Involucres | mostly campanulate, cylindric, turbinate, or hemispheric or broader. |
cylindric to obconic, 1.5–3 mm diam. (larger in fruit). |
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| Receptacles | mostly flat to convex, paleate (paleae often stramineous to white with red-brown to purplish striae, orbiculate to oblong or linear, ± flat to slightly cupped). |
slightly convex, paleate; paleae linear to subulate, scarious. |
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| Ray florets | 0 or 1–21+ (more in cultivars), pistillate and fertile, or neuter, or styliferous and sterile; corollas usually yellow to orange (sometimes bicolored: yellow to orange with brown, red-brown, or purple), sometimes cyanic or white. |
3–6, pistillate, fertile; corollas yellow. |
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| Disc florets | 3–150+, usually bisexual, fertile (functionally staminate in Dicranocarpus); corollas usually yellow to orange, sometimes brown, red-brown, or purple, tubes usually shorter than, sometimes equaling funnelform throats (longer than throats in some Thelesperma spp.), lobes (3–)5, ± deltate to lance-ovate; (staminal filaments hairy in Cosmos) anther thecae pale or dark; stigmatic papillae in 2 lines. |
3–4+, functionally staminate; corollas yellow, tubes shorter than throats, lobes 5, triangular. |
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| Phyllaries | persistent, 3–34+ in ± 2 series (usually distinct, notably connate in Thelesperma), usually ± membranous, margins usually ± scarious. |
persistent, 3–6 in 1(–2) series, lance-ovate to lanceolate, scarious, brown-nerved. |
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| Calyculi | usually of 3–8(–21+) bractlets or bracts (usually ± herbaceous, usually shorter than phyllaries and/or reflexed, sometimes ± foliaceous and surpassing phyllaries). |
of (0–)1–3+ erect, linear to subulate, herbaceous bractlets. |
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| Heads | radiate or discoid, borne singly or in ± corymbiform, cymiform, or paniculiform arrays. |
radiate, borne singly or (2–4) in cymiform arrays. |
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| Cypselae | usually either obcompressed to obflattened, usually cuneate, linear, oblong, orbiculate, or ovate (bodies unequally 3–4-angled, margins often winged), or (all or inner) ± equally 4-angled and linear-fusiform, sometimes ± beaked, faces glabrous or hairy (often striate, tuberculate, or papillate); pappi 0, or persistent, of (1–)2–4(–8), usually ± barbellate awns or scales, rarely coroniform or of 1–2, smooth to ciliate or barbed awns or scales. |
obcompressed, oblong to linear, ± ribbed, smooth or tuberculate, glabrous or densely papillate; pappi persistent, of 2 spreading to recurved, smooth or proximally papillate awns. |
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| x | = 10. |
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Asteraceae (tribe Heliantheae) subtribe Coreopsidinae |
Dicranocarpus |
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| Distribution | Mostly subtropical and warm-temperate New World and Old World |
sw United States; n Mexico |
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| Discussion | Genera 20, species 374 (7 genera, 69 species in the flora). The circumscription of Coreopsidinae followed here (H. Robinson 1981) is a bit narrower than the traditional one. The subtribe is remarkable among Heliantheae for having a relatively high number of species native in subtropical and tropical Old World, especially Africa. Distinctions among some genera (e.g., Bidens, Coreopsis, and Cosmos) are often subtle. In Coreopsidinae, each involucre is subtended by a calyculus of more or less herbaceous (sometimes leaflike) bractlets or bracts (sometimes surpassing the phyllaries). In keys and descriptions here, shapes, heights, and diameters given for involucres are based on the phyllaries collectively (exclusive of calyculi) at flowering; the involucres are sometimes notably larger in fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 1. Etymology: Greek di -, two, kranos, skull or helmet, and karpos, fruit; probably alluding to the “two-horned” cypselae (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| Synonyms | subtribe Coreopsideae, subtribe Petrobiinae | |||||||||||||||||||||||||||||
| Name authority | Lessing: Linnaea 5: 153. (1830) | A. Gray: Mem. Amer. Acad. Arts, n. s. 5: 322. (1854) | ||||||||||||||||||||||||||||
| Source | FNA vol. 21, p. 183. | FNA vol. 21, p. 219. | ||||||||||||||||||||||||||||
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