Asteraceae (tribe Heliantheae) subtribe Coreopsidinae(synonym of Asteraceae tribe Heliantheae subtribe Coreopsidinae) |
Cosmos |
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cosmos |
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| Habit | Annuals, perennials, subshrubs, or shrubs, 10–400 cm (sometimes rhizomatous or with cormiform bases, stoloniferous in Coreopsis auriculata). | Annuals [perennials or subshrubs], 30–250 cm. | ||||||||||||||||||||||||||||||||||||||||
| Stems | usually 1, erect or ascending, branched distally or ± throughout. |
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| Leaves | usually cauline (sometimes mostly basal); usually mostly opposite (distal sometimes alternate), rarely whorled; petiolate or sessile; blades (often pinnately or palmately lobed, sometimes compound), or lobes or leaflets, mostly deltate, elliptic, filiform, lanceolate, linear, oblanceolate, or ovate, ultimate margins entire or toothed, faces usually glabrous or glabrate, sometimes hairy (rarely, if ever, gland-dotted). |
mostly cauline; opposite; petiolate or sessile; blades usually 1–3-pinnately lobed [undivided], ultimate margins usually entire, faces usually glabrous, sometimes glabrate, hispid, puberulent, or scabridulous. |
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| Involucres | mostly campanulate, cylindric, turbinate, or hemispheric or broader. |
hemispheric or subhemispheric [cylindric], 3–15 mm diam. |
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| Receptacles | mostly flat to convex, paleate (paleae often stramineous to white with red-brown to purplish striae, orbiculate to oblong or linear, ± flat to slightly cupped). |
flat, paleate; paleae falling, linear, flat or slightly concave-convex, scarious (entire). |
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| Ray florets | 0 or 1–21+ (more in cultivars), pistillate and fertile, or neuter, or styliferous and sterile; corollas usually yellow to orange (sometimes bicolored: yellow to orange with brown, red-brown, or purple), sometimes cyanic or white. |
[0, 5] 8 (more in “double” cultivars), neuter; corollas white to pink or purple, or yellow to red-orange. |
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| Disc florets | 3–150+, usually bisexual, fertile (functionally staminate in Dicranocarpus); corollas usually yellow to orange, sometimes brown, red-brown, or purple, tubes usually shorter than, sometimes equaling funnelform throats (longer than throats in some Thelesperma spp.), lobes (3–)5, ± deltate to lance-ovate; (staminal filaments hairy in Cosmos) anther thecae pale or dark; stigmatic papillae in 2 lines. |
10–20[–80+], bisexual, fertile; corollas yellow [orange] (at least distally), tubes shorter than funnelform throats, lobes 5, ± deltate (staminal filaments hairy near anthers; style branches linear, flattened, thicker distally, hirtellous, appendages relatively slender). |
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| Phyllaries | persistent, 3–34+ in ± 2 series (usually distinct, notably connate in Thelesperma), usually ± membranous, margins usually ± scarious. |
persistent, [5–]8 in ± 2 series, distinct, lanceolate, lance-oblong, lance-ovate, or oblong, ± equal, membranous or herbaceous, margins ± scarious. |
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| Calyculi | usually of 3–8(–21+) bractlets or bracts (usually ± herbaceous, usually shorter than phyllaries and/or reflexed, sometimes ± foliaceous and surpassing phyllaries). |
of [5–]8 basally connate, ± linear to subulate, herbaceous (striate) bractlets. |
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| Heads | radiate or discoid, borne singly or in ± corymbiform, cymiform, or paniculiform arrays. |
radiate, borne singly or in corymbiform arrays. |
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| Cypselae | usually either obcompressed to obflattened, usually cuneate, linear, oblong, orbiculate, or ovate (bodies unequally 3–4-angled, margins often winged), or (all or inner) ± equally 4-angled and linear-fusiform, sometimes ± beaked, faces glabrous or hairy (often striate, tuberculate, or papillate); pappi 0, or persistent, of (1–)2–4(–8), usually ± barbellate awns or scales, rarely coroniform or of 1–2, smooth to ciliate or barbed awns or scales. |
(dark brown or black) relatively slender, quadrangular-cylindric or -fusiform [outer somewhat obcompressed], sometimes slightly arcuate, attenuate-beaked, not winged [winged], faces glabrous or hispid to scabridulous or ± setose, sometimes papillate, usually with 1 groove; pappi persistent [falling], of 2–4[–8] retrorsely [antrorsely] barbed awns, sometimes 0. |
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| x | = 12. |
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Asteraceae (tribe Heliantheae) subtribe Coreopsidinae |
Cosmos |
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| Distribution | Mostly subtropical and warm-temperate New World and Old World |
Tropical and subtropical America; especially Mexico; widely introduced elsewhere |
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| Discussion | Genera 20, species 374 (7 genera, 69 species in the flora). The circumscription of Coreopsidinae followed here (H. Robinson 1981) is a bit narrower than the traditional one. The subtribe is remarkable among Heliantheae for having a relatively high number of species native in subtropical and tropical Old World, especially Africa. Distinctions among some genera (e.g., Bidens, Coreopsis, and Cosmos) are often subtle. In Coreopsidinae, each involucre is subtended by a calyculus of more or less herbaceous (sometimes leaflike) bractlets or bracts (sometimes surpassing the phyllaries). In keys and descriptions here, shapes, heights, and diameters given for involucres are based on the phyllaries collectively (exclusive of calyculi) at flowering; the involucres are sometimes notably larger in fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 26 (4 in the flora). Etymology: Greek kosmos, harmoniously ordered universe, or kosmo, ornament (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| Key |
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| Synonyms | subtribe Coreopsideae, subtribe Petrobiinae | |||||||||||||||||||||||||||||||||||||||||
| Name authority | Lessing: Linnaea 5: 153. (1830) | Cavanilles: Icon. 1: 9, plate 14. (1791) | ||||||||||||||||||||||||||||||||||||||||
| Source | FNA vol. 21, p. 183. | FNA vol. 21, p. 203. | ||||||||||||||||||||||||||||||||||||||||
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