Asteraceae (tribe Heliantheae) subtribe Coreopsidinae(synonym of Asteraceae tribe Heliantheae subtribe Coreopsidinae) |
Coreopsis |
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coreopsis, tick-seed |
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| Habit | Annuals, perennials, subshrubs, or shrubs, 10–400 cm (sometimes rhizomatous or with cormiform bases, stoloniferous in Coreopsis auriculata). | Annuals, perennials, subshrubs, or shrubs, 10–80(–200+) cm (often rhizomatous or with cormiform bases, stoloniferous in C. auriculata). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Stems | usually 1, erect, branched distally or ± throughout. |
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| Leaves | usually cauline (sometimes mostly basal); usually mostly opposite (distal sometimes alternate), rarely whorled; petiolate or sessile; blades (often pinnately or palmately lobed, sometimes compound), or lobes or leaflets, mostly deltate, elliptic, filiform, lanceolate, linear, oblanceolate, or ovate, ultimate margins entire or toothed, faces usually glabrous or glabrate, sometimes hairy (rarely, if ever, gland-dotted). |
basal, basal and cauline, or cauline; opposite or alternate or both; petiolate or sessile; blades simple and entire (dentate in C. latifolia), or ± pinnately or pedately lobed, faces glabrous or hairy. |
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| Involucres | mostly campanulate, cylindric, turbinate, or hemispheric or broader. |
± globose to cylindric, 4–25+ mm diam. |
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| Receptacles | mostly flat to convex, paleate (paleae often stramineous to white with red-brown to purplish striae, orbiculate to oblong or linear, ± flat to slightly cupped). |
flat to convex, paleate; paleae falling, ovate to linear or subulate, ± flat, ± scarious (not adnate to and not falling with cypselae except in C. bigelovii). |
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| Ray florets | 0 or 1–21+ (more in cultivars), pistillate and fertile, or neuter, or styliferous and sterile; corollas usually yellow to orange (sometimes bicolored: yellow to orange with brown, red-brown, or purple), sometimes cyanic or white. |
mostly (5–)8(–12+, more in “double” cultivars), neuter, or styliferous and sterile, or pistillate and fertile; corollas usually yellow, sometimes red-brown to purple proximally, sometimes wholly purple or pink to white. |
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| Disc florets | 3–150+, usually bisexual, fertile (functionally staminate in Dicranocarpus); corollas usually yellow to orange, sometimes brown, red-brown, or purple, tubes usually shorter than, sometimes equaling funnelform throats (longer than throats in some Thelesperma spp.), lobes (3–)5, ± deltate to lance-ovate; (staminal filaments hairy in Cosmos) anther thecae pale or dark; stigmatic papillae in 2 lines. |
8–150+, bisexual, fertile; corollas usually yellow, sometimes red-brown to purple at tips or throughout, tubes equaling or shorter than narrowly funnelform throats, lobes 4 or 5 (adaxial sinus seldom deeper than others). |
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| Phyllaries | persistent, 3–34+ in ± 2 series (usually distinct, notably connate in Thelesperma), usually ± membranous, margins usually ± scarious. |
usually ± 8 in ± 2 series (usually distinct, rarely connate ± 1/10 their lengths, mostly oblong to linear, ± membranous, margins ± scarious). |
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| Calyculi | usually of 3–8(–21+) bractlets or bracts (usually ± herbaceous, usually shorter than phyllaries and/or reflexed, sometimes ± foliaceous and surpassing phyllaries). |
of (3–)8+, distinct, ± herbaceous bractlets. |
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| Heads | radiate or discoid, borne singly or in ± corymbiform, cymiform, or paniculiform arrays. |
radiate, borne singly or in open, ± corymbiform arrays. |
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| Cypselae | usually either obcompressed to obflattened, usually cuneate, linear, oblong, orbiculate, or ovate (bodies unequally 3–4-angled, margins often winged), or (all or inner) ± equally 4-angled and linear-fusiform, sometimes ± beaked, faces glabrous or hairy (often striate, tuberculate, or papillate); pappi 0, or persistent, of (1–)2–4(–8), usually ± barbellate awns or scales, rarely coroniform or of 1–2, smooth to ciliate or barbed awns or scales. |
obcompressed, ± orbiculate to ovate, oblong, or linear, usually thin-margined or winged, wings membranous to chartaceous or corky, entire or lobed to toothed, sometimes ciliolate; faces smooth or ± papillate to tuberculate; pappi 0, or persistent, of 2 bristly cusps or scales (sometimes pappi 0 and shoulders of cypsela wings ± bristly, pappus-like). |
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| x | = 14. |
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Asteraceae (tribe Heliantheae) subtribe Coreopsidinae |
Coreopsis |
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| Distribution | Mostly subtropical and warm-temperate New World and Old World |
Mostly temperate North America; also tropical New World and Old World Early leaves of some coreopsises often differ from later leaves on individual plants |
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| Discussion | Genera 20, species 374 (7 genera, 69 species in the flora). The circumscription of Coreopsidinae followed here (H. Robinson 1981) is a bit narrower than the traditional one. The subtribe is remarkable among Heliantheae for having a relatively high number of species native in subtropical and tropical Old World, especially Africa. Distinctions among some genera (e.g., Bidens, Coreopsis, and Cosmos) are often subtle. In Coreopsidinae, each involucre is subtended by a calyculus of more or less herbaceous (sometimes leaflike) bractlets or bracts (sometimes surpassing the phyllaries). In keys and descriptions here, shapes, heights, and diameters given for involucres are based on the phyllaries collectively (exclusive of calyculi) at flowering; the involucres are sometimes notably larger in fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 35 (28 in the flora). Early leaves of some coreopsises often differ from later leaves on individual plants. In such plants, early (proximal) leaves may be 1–3-pinnately or pedately lobed and 12–25+ cm long with 9–15+ orbiculate to lanceolate, linear, or filiform lobes and contrast markedly with later (distal) leaves 1–3 cm that are undivided or have 3–5+ lanceolate to linear or filiform lobes. Cultivars (often “doubles” with multiple series of “ray” florets) derived from Coreopsis auriculata, C. grandiflora, C. lanceolata, and C. tinctoria are grown in public and residential gardens and are grown commercially for cut flowers. Etymology: Greek korios, bedbug, and -opsis, resembling, alluding to cypselae of original species (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| Synonyms | subtribe Coreopsideae, subtribe Petrobiinae | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Name authority | Lessing: Linnaea 5: 153. (1830) | Linnaeus: Sp. Pl. 2: 907. (1753): Gen. Pl. ed. 5, 388. (1754) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
| Source | FNA vol. 21, p. 183. | FNA vol. 21, p. 185. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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