Asteraceae (tribe Heliantheae) subtribe Coreopsidinae(synonym of Asteraceae tribe Heliantheae subtribe Coreopsidinae) |
Coreocarpus |
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coreocarpus |
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| Habit | Annuals, perennials, subshrubs, or shrubs, 10–400 cm (sometimes rhizomatous or with cormiform bases, stoloniferous in Coreopsis auriculata). | Perennials [subshrubs], 10–70(–120+) cm. | ||||||||||||||||||||||||||||
| Stems | 1, erect [sprawling], branched throughout. |
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| Leaves | usually cauline (sometimes mostly basal); usually mostly opposite (distal sometimes alternate), rarely whorled; petiolate or sessile; blades (often pinnately or palmately lobed, sometimes compound), or lobes or leaflets, mostly deltate, elliptic, filiform, lanceolate, linear, oblanceolate, or ovate, ultimate margins entire or toothed, faces usually glabrous or glabrate, sometimes hairy (rarely, if ever, gland-dotted). |
cauline; opposite; obscurely petiolate; blades 1–2-pinnately or -pedately lobed (ultimate lobes narrowly lanceolate to filiform [lanceolate to ovate]), ultimate margins entire, faces usually glabrous, rarely hairy [pilose]. |
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| Involucres | mostly campanulate, cylindric, turbinate, or hemispheric or broader. |
± campanulate to turbinate, 3–5+ mm diam. |
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| Receptacles | mostly flat to convex, paleate (paleae often stramineous to white with red-brown to purplish striae, orbiculate to oblong or linear, ± flat to slightly cupped). |
flat to convex, paleate; paleae falling, (stramineous with red-brown striae) lance-linear to lanceolate, membranous to scarious [adnate to cypselae]. |
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| Ray florets | 0 or 1–21+ (more in cultivars), pistillate and fertile, or neuter, or styliferous and sterile; corollas usually yellow to orange (sometimes bicolored: yellow to orange with brown, red-brown, or purple), sometimes cyanic or white. |
0, or [1–]5–8, pistillate, fertile [styliferous and sterile or neuter]; corollas [orange] yellow to pale yellow or white [purplish]. |
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| Disc florets | 3–150+, usually bisexual, fertile (functionally staminate in Dicranocarpus); corollas usually yellow to orange, sometimes brown, red-brown, or purple, tubes usually shorter than, sometimes equaling funnelform throats (longer than throats in some Thelesperma spp.), lobes (3–)5, ± deltate to lance-ovate; (staminal filaments hairy in Cosmos) anther thecae pale or dark; stigmatic papillae in 2 lines. |
[5–]12–25[–30+], bisexual, fertile; corollas yellow with red-brown nerves, tubes shorter than narrowly funnelform throats, lobes 5, ± deltate. |
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| Phyllaries | persistent, 3–34+ in ± 2 series (usually distinct, notably connate in Thelesperma), usually ± membranous, margins usually ± scarious. |
persistent, 5–8[–13+] in ± 2 series, distinct, (green to stramineous with red-brown to purplish nerves) ovate to oblong, ± equal, membranous, margins scarious. |
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| Calyculi | usually of 3–8(–21+) bractlets or bracts (usually ± herbaceous, usually shorter than phyllaries and/or reflexed, sometimes ± foliaceous and surpassing phyllaries). |
of (0–)1–3+ linear to subulate, herbaceous bractlets. |
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| Heads | radiate or discoid, borne singly or in ± corymbiform, cymiform, or paniculiform arrays. |
usually radiate, sometimes discoid, in open, corymbiform to paniculiform arrays [borne singly]. |
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| Cypselae | usually either obcompressed to obflattened, usually cuneate, linear, oblong, orbiculate, or ovate (bodies unequally 3–4-angled, margins often winged), or (all or inner) ± equally 4-angled and linear-fusiform, sometimes ± beaked, faces glabrous or hairy (often striate, tuberculate, or papillate); pappi 0, or persistent, of (1–)2–4(–8), usually ± barbellate awns or scales, rarely coroniform or of 1–2, smooth to ciliate or barbed awns or scales. |
(blackish or red-brown to stramineous) obcompressed, ± oblanceolate, faces smooth or ± granular-papillate, margins ± winged, wings ± corky, ± pectinate [entire]; pappi 0, or persistent, of 1–2 retrorsely [antrorsely] barbellate awns. |
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| x | = 12. |
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Asteraceae (tribe Heliantheae) subtribe Coreopsidinae |
Coreocarpus |
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| Distribution | Mostly subtropical and warm-temperate New World and Old World |
sw United States; nw Mexico |
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| Discussion | Genera 20, species 374 (7 genera, 69 species in the flora). The circumscription of Coreopsidinae followed here (H. Robinson 1981) is a bit narrower than the traditional one. The subtribe is remarkable among Heliantheae for having a relatively high number of species native in subtropical and tropical Old World, especially Africa. Distinctions among some genera (e.g., Bidens, Coreopsis, and Cosmos) are often subtle. In Coreopsidinae, each involucre is subtended by a calyculus of more or less herbaceous (sometimes leaflike) bractlets or bracts (sometimes surpassing the phyllaries). In keys and descriptions here, shapes, heights, and diameters given for involucres are based on the phyllaries collectively (exclusive of calyculi) at flowering; the involucres are sometimes notably larger in fruit. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 9 (1 in the flora). Etymology: Greek koreos, bug, and karpos, fruit, alluding to pectinately winged cypselae of original species (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| Key |
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| Synonyms | subtribe Coreopsideae, subtribe Petrobiinae | |||||||||||||||||||||||||||||
| Name authority | Lessing: Linnaea 5: 153. (1830) | Bentham: Bot. Voy. Sulphur 28, plate 16. (1844) | ||||||||||||||||||||||||||||
| Source | FNA vol. 21, p. 183. | FNA vol. 21, p. 198. | ||||||||||||||||||||||||||||
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