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brightgreen spleenwort, doradille verte, green spleenwort

adulterated spleenwort, corrupt spleenwort, ladder spleenwort

Roots

not proliferous.

not proliferous.

Stems

short-creeping or ascending, frequently branched;

scales dark reddish brown to blackish throughout, narrowly deltate, 2–4 × 0.2–0.4 mm, margins entire to undulate or with widely spaced shallow teeth.

short-creeping, mainly unbranched;

scales black or with narrow pale borders, narrowly lanceolate, 1.5–3 × 0.2–0.4 mm, margins entire.

Leaves

monomorphic.

monomorphic.

Petiole

reddish brown at base, green distally, lustrous, 1–5(–6) cm, 1/4–1/2(–1) times length of blade;

indument of dark reddish brown to black, narrowly deltate scales grading into glandular hairs.

dark reddish brown throughout, 1–4 mm;

indument of black linear scales at base.

Blade

linear, 1-pinnate throughout, 2–13 × 0.6–1.2 cm, thin, glabrous or with sparse minute hairs;

base slightly tapering or truncate;

apex acute, not rooting.

linear, 1-pinnate, 2.5–14 × 0.5–1.2 cm, thick (open habitat) to herbaceous (shaded, moist habitat), essentially glabrous;

base somewhat tapered;

apex obtuse, not rooting.

Pinnae

in 6–21 pairs, deltate to rhombic;

medial pinnae 5–6 × 4–5 mm;

base obtuse and often inequilateral;

distal margins crenate;

apex rounded to acute.

in 10–30 pairs, ovate to rhombic to ovate-oblong, 2.5–11 × 2–6 mm;

base truncate to shortly acute;

margins shallowly crenate (shade forms) to essentially entire (exposed forms);

apex obtuse, broadly rounded.

Veins

free, evident.

free, evident to obscure.

Sori

2–4 pairs per pinna, on both basiscopic and acroscopic sides.

1–3 pairs per pinna on both basiscopic and acroscopic sides.

Spores

64 per sporangium.

64 per sporangium.

Rachis

green throughout, dull, glabrous or with scattered hairs as on petioles.

reddish brown in proximal 1/2–4/5, green distally, lustrous, glabrous.

2n

= 72.

= 144.

Asplenium trichomanes-ramosum

Asplenium adulterinum

Habitat Limestone and other basic rocks Crevices in limestone
Elevation 0–4000 m (0–13100 ft) 1250 m (4100 ft)
Distribution
from FNA
AK; CA; CO; ID; ME; MI; MT; NV; NY; OR; SD; UT; VT; WA; WI; WY; AB; BC; NB; NF; NS; NT; ON; PE; QC; YT; Greenland
[WildflowerSearch map]
from FNA
BC; Europe
[BONAP county map]
Discussion

Hybridization between Asplenium trichomanes-ramosum and A. trichomanes produces the fertile allotetraploid A. adulterinum, which occurs on Vancouver Island.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

In North America Asplenium adulterinum is known to occur on Vancouver Island, British Columbia, where only the fertile allotetraploids are known. It is likely to occur in areas where the two parents, A. trichomanes and A. trichomanes-ramosum, grow together. The genetics of the American plants should be compared with that of the European, among which two nothosubspecies occur (F. Mokry et al. 1986).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 2. FNA vol. 2.
Parent taxa Aspleniaceae > Asplenium Aspleniaceae > Asplenium
Sibling taxa
A. abscissum, A. adiantum-nigrum, A. adulterinum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
A. abscissum, A. adiantum-nigrum, A. auritum, A. bradleyi, A. cristatum, A. dalhousiae, A. ebenoides, A. exiguum, A. heterochroum, A. monanthes, A. montanum, A. palmeri, A. pinnatifidum, A. platyneuron, A. plenum, A. pumilum, A. resiliens, A. rhizophyllum, A. ruta-muraria, A. scolopendrium, A. septentrionale, A. serratum, A. trichomanes, A. trichomanes-dentatum, A. trichomanes-ramosum, A. verecundum, A. vespertinum, A. ×biscayneanum, A. ×curtissii, A. ×heteroresiliens
Synonyms A. viride
Name authority Linnaeus: Sp. Pl. 2: 1082. (1753) J. Milde: Höh. Sporenpfl. Deutschl. 40. (1865)
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